Olivoidea, Latreille, 1825

Superfamily Olivoidea View in CoL Family Olividae

Genus Agaronia Gray, 1839

Agaronia sterica new species ( Figs. 13-14 View Figure 13 View Figure 14 ) https://zoobank.org/ 31899908-C9BC-451E-AAF2-F6D62F828E37

Agaronia steeriae View in CoL : Morretes, 1949: 101; Rios, 1970: 102 (pl. 32); Teso & Pastorino, 2011: 24 (in syn of O. carcellesi ) (non Reeve, 1850).

Olivancillaria steeriae View in CoL : Rios, 1975: 110 (pl. 32, fig. 465), 1985: 110 (in syn of O. vesica View in CoL ), 1994: 142 (in syn of O. vesica View in CoL ), 2009: 270 (in syn of O. vesica View in CoL ); Oliveira et al., 1981: 223; Teso & Pastorino, 2011: 24 (in syn of O. carcellesi ) (non Reeve, 1850).

Olivancillaria cf. carcellesi View in CoL : Teso & Pastorino, 2011: 25 (fig. 15G, H, J) (non Klappenbach, 1965).

Types: Holotype MZSP 166100, ♀. Paratypes: MZSP 36498, 1♂ from type locality. BRAZIL. Espírito Santo; Vitória, Praia da Guarderia View in CoL ( Praia Comprida View in CoL ), 20°18′24″S 40°17′07″W, MZSP 51042, 3 shells (Castro col., o.t., xii.1974); Guarapari, off, 20°54′12″S 40°07′55″W, 50-60 m, MZSP 69465, 2 spm (Femorale #18772, o.t., vii.2005). Rio de Janeiro; Campos dos Goytacazes, off Cabo de São Tomé, 22°05′02″S 40°45′10″W, 30-40 m, MZSP 73374, 3 shells (Femorale, o.t., ix.2000); Cabo Frio, off, 22°55′30″S 41°57′44″W, 30-40 m, MZSP 68586, 15 spm (Femorale #21803, o.t., ix.2005).

Type locality: BRAZIL. Espírito Santo ; Meaípe, off Meaípe, 20°46′32″S 40°31′35″W, 20-25 m [o.t., x.1993].

Diagnosis: SE Brazilian species generally with 35-40 mm, slender. Color uniform brownish grey to spotted brown on beige base. Spire short, bluntly angled. Aperture narrow, with narrow callus; inferior half of inner lip weakly folded obliquely. Radula with rachidian wide, with samesized central cusps.

Description: Shell ( Figs. 13 View Figure 13 , 14 View Figure 14 A-B) up to 50 mm. Fusiform, lateralized; ~2.0-2.2 times longer than wide; spire~20% of total length, last whorl ~93% of total length. Spire angle ~70°. Spire up to 4 plane whorls; suture pane, only seen by narrow furrow. Protoconch of ~2 smooth, glossy whorls, ~ 1.8 mm, white ( Fig. 13E View Figure 13 ); profile blunt ( Fig. 13 View Figure 13 G-K), slightly pointed ( Fig. 13 View Figure 13 A-D, L-M) or mamillated ( Figs. 13 View Figure 13 O-P, 14A-B). Shell surface smooth, glossy, weak growth lines. Color siphonal slope always brown, with narrow ( Fig. 13J, K View Figure 13 ), to wide middle spiral white band ( Fig. 13H, P View Figure 13 ), sometimes absent ( Fig. 13D View Figure 13 ); remaining body whorl variating from uniform gray ( Fig. 13 View Figure 13 B-D), predominantly gray with some beige ( Fig. 13L, N View Figure 13 ), basic light beige with brown drizzle denser ( Figs. 13J, K, P View Figure 13 , 14B View Figure 14 ), or sparser ( Fig. 13H View Figure 13 ); spire variating from brown ( Fig. 13D View Figure 13 ), beige ( Fig. 13 View Figure 13 G-K), light beige ( Fig. 13P View Figure 13 ), to bicolor ( Figs. 13N View Figure 13 , 14B View Figure 14 ). Aperture elongated, ~4-times longer than wide; superior end continuing with sutural narrow furrow. Canal widely opened, of ~40% of last whorl width. Outer lip simply arched. Inner lip white, weakly concave, almost straight; callus relatively thick in larger specimens ( Figs. 13A, B, O View Figure 13 , 14A View Figure 14 ), located in superi- or region, sometimes covering part of spire; sometimes white ( Fig. 13O View Figure 13 ), sometimes with brown spot ( Fig. 13S View Figure 13 ); callus narrowing in middle level of inner lip; inferior half of inner lip with set of ~10 oblique narrow, relatively uniform folds, located close from each other ( Fig. 13C, M View Figure 13 ); superior fold usually larger than others,groove as inferior limit of this set of folds, ending in anterior blunt beak (Fi gs. 13B, C, G, I, L, M, O, 14A). No umbilicus. No operculum.

Head-foot entirely retracted inside shell ( Fig. 13A, G, I View Figure 13 ). Head small, marked by small pair of cephalic flaps ( Fig. 14E View Figure 14 : cp); eyes and tentacles absent. Foot very wide and ample ( Fig. 14G View Figure 14 : ft); anterior edge covered by well-developed propodium ( Fig. 14E, G View Figure 14 : pr), laterally expanding in pair of small, pointed lateral projections ( Fig. 14G View Figure 14 : fl). Mandle edge relatively thick, unpigmented ( Fig. 14C, E View Figure 14 : mb); pallial tentacle ( Fig. 14E, F View Figure 14 : pt) located in posterior end of mantle edge; basal region broad, tapering gradually up to pointed tip. Pallial cavity rather triangular, lateral positioned, gill ( Fig. 14D View Figure 14 : gi) and osphradium located in left side; wide pallial gonoducts in right side). Rhynchostome pore-like, located between and ventral to cephalic flaps.

Radula ( Fig. 13 View Figure 13 H-I) rachidian ~40% of radular width; ~3 times wider than long; base chevron-like, posterior arched, concave; cutting edge with 3 central, tall cusps, divergently positioned, median cusp slightly smaller than lateral cusps;single very small cusps in base of lateral cusps; remaining marginal rachidian third as flattened, simple weakly arched rod. Lateral teeth with wide base (~50% of rachidian width), abruptly tapering up to hooklike terminal tip, tuned inside.

Pallial oviduct occupying ~ ⅓ of pallial cavity, round- ed; displacing rectum and anus to left, anus aperture away from right corner ( Fig. 14D View Figure 14 : an). Bursa copulatrix spheric, located in middle level of pallial oviduct posterior edge ( Fig. 14D View Figure 14 : bc); bursa duct (bd) wide, running transversely along oviduct ventral wall dividing oviduct approximately in half, up to region close to anus. Penis filiform, very elongated ( Fig. 14E View Figure 14 : pe), inserted at some distance posterior and at right from cephalic flaps, tapering very gradually; tip bluntly pointed; penis duct entirely closed (tubular), opening at tip.

Etymology: The specific epithet is the nickname of the species when the Brazilian population was called "steeriae ", a joking related to the name hysterical, related to the confusion with the African species, and the lability of shell colors.

Distribution: Espírito Santo and Rio de Janeiro coasts.