Benthophilus granulosus Kessler, 1877

Zarei, Fatah, Esmaeili, Hamid Reza, Abbasi, Keyvan, Kovačić, Marcelo, Schliewen, Ulrich K. & Stepien, Carol A., 2022, Gobies (Teleostei: Gobiidae) of the oldest and deepest Caspian Sea sub-basin: an evidence-based annotated checklist and a key for species identification, Zootaxa 5190 (2), pp. 151-193 : 166

publication ID	https://doi.org/10.11646/zootaxa.5190.2.1
publication LSID	lsid:zoobank.org:pub:92BFDA7A-28C0-4816-8A91-CFB842C95E96
DOI	https://doi.org/10.5281/zenodo.7120094
persistent identifier	https://treatment.plazi.org/id/4154F53A-8A13-FF95-FF72-ACD8FDCF4E73
treatment provided by	Plazi
scientific name	Benthophilus granulosus Kessler, 1877
status

Treatment

8. Benthophilus granulosus Kessler, 1877 View in CoL ( Fig. 10 View FIGURE 10 ), Granular Tadpole Goby

Benthophilus granulosus Kessler, 1877: 57 View in CoL , fig. 14; type locality: Baku Bay , Caspian Sea, Azerbaijan; lectotype: ZISP [= ZIN] 2242, syntypes: (7 + 1) BMNH 1897.7 .5.14 [ex ZIN] (1), ZIN 10898 (1, damaged).

Etymology: Latin, granulatus = covered or composed of grains, refers to the granular appearance of the skin.

Distribution and habitat: Coastal waters in all three sub-basins of the Caspian Sea ( Fig. 11A View FIGURE 11 ), also enters river deltas. Eurybathic, during the warm season, it inhabits shallow coastal waters ( 0.5–20 m), then migrates to deeper places (down to 60–70 m) in the winter ( Boldyrev & Bogutskaya 2007). Marked for Iran on the minimap in Pinchuk & Miller ( 2004x), but according to Boldyrev & Bogutskaya (2007) lacked Iranian records, but now has been confirmed in Iran by Abbasi (2017) and by the present study’s examined material.

Material examined: ZM-CBSU S006, 1, off Anzali & ZM-CBSU S023, 2, off Chaboksar , Gilan Province, Iran .

IUCN: LC ( Freyhof & Kottelat 2008c).

Kingdom	Animalia
Phylum	Chordata
Class	Actinopterygii
Order	Perciformes
Family	Gobiidae
Genus	Benthophilus