Bollmannia nubila Schwarzhans, 2010

Bollmannia nubila Schwarzhans, 2010

Figure 16a–b View Fig

2010 Bollmannia nubila —Schwarzhans: pl. 101, Fig. 1 View Fig .

2014 Gobiidae , genera and species indet. 1—Aguilera et al.: Fig. 12 View Fig /5–7.

Material 9 specimens: holotype (refigured from Schwarzhans, 2010) from the Langhian of Dingden , northern Germany, North Sea Basin ( SMF PO 83052 ). Aquitanian to Burdigalian: Brazil, Pirabas FM , 8 specimens (1 specimen refigured from Aguilera et al., 2014 as Gobiidae indet.) (MPEG- 1820-V).

Description Otolith size up to 2 mm in length. OL:OH = 1.05–1.1; OH:OT = 3.0–3.2. Otolith shape nearly rectangular, predorsal angle high, nearly as high as postdorsal angle, postdorsal projection blunt, not longer than postventral angle, not bent outward. Ventral rim straight, horizontal, smooth. Anterior and posterior rims nearly vertical; preventral angle blunt, nearly orthogonal; posterior rim with broad, shallow indentation at its middle. All rims smooth, dorsal rim occasionally crenulated.

Inner face nearly flat. Sulcus sole-shaped but very narrow with low ostial lobe, slightly deepened, inclined at 10–15°. OL:SuL = 1.65–1.85. Subcaudal iugum long, relatively narrow. Dorsal depression wide, indistinct. Ventral furrow distinct, close to ventral rim of otolith. Outer face distinctly convex, strongest inframedian, relatively smooth with few short furrows on dorsal field.

Discussion Bollmannia nubila is the earliest record of the genus and now known from two rather extreme geographic positions: the unique type from the North Sea Basin and the now-assigned referred specimens from Brazil. Bollmannia nubila is readily distinguished from its congeners by the very narrow sulcus, the relatively narrow subcaudal iugum, and the high predorsal angle. It resembles B. eigenmannorum . It is therefore possible that B. nubila is positioned relatively closely to the origin of the genus and thus combines characteristics of the B. boqueronensis plexus and the B. eigenmannorum plexus.