Anomala spiniforceps Zhao, Fujioka & Zorn, 2025

Anomala spiniforceps Zhao, Fujioka & Zorn View in CoL , sp. nov.

[Chinese vernacular name: 刺ĩ异丽aek]

( Figs 3A–B View FIGURE 3 , 10C View FIGURE 10 , 16A–D View FIGURE 16 )

Type material ( 3♂♂, 4♀♀). Holotype: ♂ ( NSMT) “China-India border: W. Siang 03. JULY. 2011 Takeshi MAEDA-leg.” Paratypes: 1♀ ( MFPC) “China-India border: W. Siang 03. JULY. 2011 Takeshi MAEDA-leg.” ; 1♂, 1♀ ( ZMPC) “China-India border: W. Siang 30. JUNE. 2011 Takeshi MAEDA-leg.” ; 1♂, 1♀ ( MFPC) “China-India border: alt. 1700m Lower Subansiri 24–27. JUN. 2015 Takeshi Maeda-leg.” ; 1♀ ( MFPC) “China-India border: alt. 1700m Lower Subansiri 12–20. AUG. 2015 Takeshi Maeda-leg.”

Description. Male. Body shape elongated ovoid, moderately convex.

Color . Generally brown to black, elytra sometimes lighter; shiny, with strong greenish luster; elytra with a yellow, broad zigzag band extending from primary costa 5 to 1; antennae reddish or yellowish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown.

Head. Greatest width/length of clypeus approximately 2.4; subtrapezoidal, lateral margins straight and moderately convergent anteriad, anterior margin slightly convex, anterior corners obsolete and broadly rounded; anterior margin weakly reflexed; surface very densely and coarsely rugopunctate. Frontoclypeal suture straight, distinct. An inverted triangular area on anterior third of frons rugopunctate, other portions with dense and large punctures, slightly sparser and smaller posteriad. Interocular distance/maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1–6. Inner margin of eye with a few moderately long setae.

Pronotum. Sides strongly convergent anteriad and weakly curved. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and large punctures. Lateral margin with several long setae.

Scutellum triangular, lateral margin moderately excurved; with dense and small punctures.

Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, moderately dense and large punctures, interstice II with a somewhat irregular secondary stria, all intervals bearing moderately dense punctures, which are slightly smaller than strial ones. Humeral protuberance moderately prominent, apical protuberance strongly prominent. Lateral carina weakly expanded from base to apical two fifths. Epipleuron with a row of sparse, short to moderately long setae from base to apical fifth.

Abdomen. Propygidium with very dense, transverse and very fine striation. Pygidium with dense and transverse striation, sparse only on the greatest tumidity; with some long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1–3 strongly carinate laterally; ventrites 2–5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated, denser laterad, frequently confluent at the lateralmost area, ventrite 6 very densely and finely rugostriolate; ventrites 2–5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin.

Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur/narrowest extremity of mesosternum approximately 6.0.

Legs. Protibia bidentate, both teeth distinct; apical tooth extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw not convex, forming a notch internobasally; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional triangular basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae.

Male genitalia. See fig. 16A–D.

Female. Sometimes with weak purple luster. Length of antennal club approximately 0.8 times as long as combined length of antennomeres 1–6. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially.Apical protibial tooth spatulate; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, basal denticle of metatarsomere 5 rather weakly developed; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw shorter than the upper one.

Measurement. Body length: 14.3–15.2 mm in males and 13.8–14.7 mm in females, greatest width: 8.1–8.4 mm in males and 8.0– 9.1 mm in females.

Differential diagnosis. The bifurcate upper branch of the left paramere of Anomala spiniforceps sp. nov. resembles that of A. flavonotata . However, the additional branch is situated near the middle in the new species (situated in the apical third in A. flavonotata ). The lower branches of both parameres have long and acute internoapical angles in A. spiniforceps sp. nov., whereas they are distinctly shorter or absent in A. flavonotata .

Etymology. The name spiniforceps is derived from the Latin words spina (spine) and forceps. It refers to the peculiar parameres, which bear prominent spine-like projections.