Anomala menyuensis Zhao, Fujioka & Zorn, 2025

Anomala menyuensis Zhao, Fujioka & Zorn View in CoL , sp. nov.

[Chinese vernacular name: mā异丽aek]

( Figs 2A–C View FIGURE 2 , 10B View FIGURE 10 , 15E–H View FIGURE 15 )

Type material ( 12♂♂, 3♀♀). Holotype: ♂ ( NHMB) “ China-India border: DIRANG vicinity, 1550± 150m 27°21′- 23′N 92°13′-16′ E L. Dembický leg.; 1-9.vi.2004 ” . Paratypes: 8♂♂, 2♀♀ ( NHMB), “ China-India border: DIRANG vicinity, 1550± 150m 27°21′-23′N 92°13′-16′ E L. Dembický leg.; 1-9.vi.2004 ” ; 2♂♂, 1♀ ( ZMPC), “ China-India border: DIRANG vicinity, 1550± 150m 27°21′-23′N 92°13′-16′ E L. Dembický leg.; 1-9.vi.2004 ” ; 1♂ ( ZMPC) “ China-India border: East Kameng , Chayangtajo circle, Laching bagang, 2200m 2024.VIII.22 ” .

Description. Male. Body shape elongated ovoid, moderately convex.

Color . Generally brown to black, the elytra sometimes lighter; shiny, with weak purplish or greenish luster, head tinged with bronze luster; the elytra with a yellow, narrow to broad zigzag band extending from primary costa 5 to 1, sometimes partly interrupted; antennae yellowish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown.

Head. Greatest width/length of clypeus approximately 2.4; subtrapezoidal, lateral margins straight and weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface with very dense, large and irregular punctures. Frontoclypeal suture straight, distinct. Anterior half of frons with dense and large punctures, gradually sparser and smaller posteriad. Interocular distance/maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1–6. Inner margin of eye with a few moderately long setae.

Pronotum. Sides strongly convergent anteriad and weakly curved. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line indistinct and broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and small punctures. Lateral margin with several long setae.

Scutellum triangular, lateral margin nearly straight; with moderately dense and minute punctures.

Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, irregular and large punctures, interstice II with a somewhat irregular secondary stria, all intervals bearing moderately dense and minute punctures, most sparse on subsutural interstice. Humeral protuberance moderately prominent, apical protuberance strongly prominent. Lateral carina weakly expanded from base to apical two fifths. Epipleuron with a row of dense, short to moderately long setae from base to apical protuberance.

Abdomen. Propygidium with moderately dense and very transverse punctures. Pygidium with dense and transverse striation, gradually breaking up into sparse and transverse punctures towards the smooth greatest tumidity; with some long setae preapically and a transverse row of long setae along posterior margin. Abdominal ventrites 1–3 strongly carinate laterally; ventrites 2–5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 with very dense and irregular punctures; ventrites 2–5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin.

Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur/narrowest extremity of mesosternum approximately 6.0.

Legs. Protibia bidentate, both teeth distinct; apical tooth acute or rounded apically, extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw not convex, forming a notch internobasally; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional rounded basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae.

Male genitalia. See fig. 15E–H.

Female. The zigzag band on elytra broad, sometimes interrupted where reaching subsutural interstice. Interocular distance/maximum head width approximately 0.7; length of antennal club approximately 0.8 times as long as combined length of antennomeres 1–6. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, subapical tooth somewhat rounded apically; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, basal denticle of metatarsomere 5 absent; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider and shorter than upper branch.

Measurement. Body length: 15.8–16.1 mm in males and 15.8–16.1 mm in females, greatest width: 9.1 mm in males and 9.0– 9.2 mm in females.

Differential diagnosis. Anomala menyuensis sp. nov. is closely related to A. flavonotata but can be reliably distinguished from this species by the left paramere, whose dorsal branch has an enlarged, rounded apex in lateral view with a distinct subapical protuberance, whereas in A. flavonotata , the apex is simply pointed. Additionally, the epipleura of A. menyuensis sp. nov. are densely covered with setae, in contrast to the sparse setation in A. flavonotata .

Remarks. We examined a specimen of A. flavonotata from Tawang (only ca. 35 km northwest of the type locality of A. menyuensis sp. nov., Dirang) exhibiting morphological consistency with specimens from Sikkim and Bhutan. Despite their similar male genitalia, the stability of A. flavonotata across its distribution range suggests that it is distinct from A. menyuensis sp. nov.

Etymology. The specific epithet menyuensis is derived from Menyu (also spelled Monyul; mā in Chinese), a historical regional name in southeastern Tibet. The species is currently known only from this region.