Anomala flavonotata Arrow, 1912

Anomala flavonotata Arrow, 1912 View in CoL

[Chinese vernacular name: ƀū异丽aek]

( Figs 1A–F View FIGURE 1 , 10A View FIGURE 10 , 14A–H View FIGURE 14 , 15A–D View FIGURE 15 , 25A, 25G–J View FIGURE 25 )

Anomala flavonotata Arrow, 1912: 73 View in CoL [original type locality: “ Sikkim: Kurseong, 5000 ft. ”, “Darjeeling”]; Arrow 1917: 212; Zorn & Bezděk 2016: 333.

Type material examined ( 1♂, 2♀♀). Lectotype (hereby designated): 1♂ ( NHMUK) “ ♂ // Type // Kurseong , Sikkim R. P. Verschraeghen. 1911-218. // Kurseong Inde Verschraeghen 1904 1907 // Anomala flavonotata, Arrow Type // NHMUK010844184” . Paralectotypes: 1♀ ( NHMUK) “ ♀ // Kurseong, Sikkim R. P. Verschraeghen. 1911- 218. // Kurseong Inde Verschraeghen 1904”; 1♀ ( MNHN) “ Kurseong Inde Verschraeghen 1904 1908 // Anomala flavonotata Arr. Déterminé par Arrow 1912 // Type // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” .

Additional material examined ( 25♂♂, 23♀♀). 1♀ ( GSPC), “N.E. INDIA Darjeeling” ; 1♂, 1♀ ( ZMPC) “N.E. INDIA Darjeeling” ; 1♂ ( GSPC) “ Umgebung Darjeeling 20.8. // anom. flavonotata , arr. Compared with type. G.J.A.” ; 1♂ ( NMBF) “ Darjeeling W.B. 2180 m, VI.1961 leg. G. Scherer // Anomala flavonotata Arr det. G. Frey, 1962” ; 2♀♀ ( MHNG) “ Kurseong, Darjeeling // Coll. R. Petrovitz ” ; 9♂♂, 8♀♀ ( MNHN) “ Kurseong R. P. Decoly 1898 // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” ; 1♂, 1♀ ( ZMPC) “ Kurseong R. P. Decoly 1898 // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” ; 1♀ ( MNHN) “ Darjeeling Inde britan. Coll. J. Clermont // MUSÉUM PARIS 1944 Coll. E. BENDERITTER” ; 1♂ ( CZPC) “ INDIA – SIKKIM west PEELING env., 2100m 18.- 20.7.1997 Jan Schneider lgt.” ; 1♂, 1♀ ( CZPC) “north India - west SIKKIM Khecheopari lake ( 1800 m.m) 4.- 10.6.1999 lgt. E. Kučera” ; 1♂ ( ZMPC) “north INDIA - west SIKKIM Khecheopari lake ( 1800 m.m) 4.- 10.6.1999 lgt. E. Kučera” ; 1♂ ( MFPC) “ INDIA - SIKKIM Khecheopalri lake 15.6– 22.6.2008 1800 m leg. E. Kučera ” ; 1♂ ( NMPC) “north INDIA - west Bengal Shingalila national park Tonglu ( 3070 m.m) 14.-19.6.99 lgt. E. Kučera ” ; 1♀ ( CZPC) “north India - west SIKKIM Shingalila national park Tonglu ( 3070 m.m) 14.- 19.6.1999 lgt. E. Kučera ” ; 1♀ ( CZPC) “ India. SIKKIM east GANTOK env., 2000-2500m FAMBONG LHO forest 8.- 15.7.1997 Jan Schneider lgt.” ; 1♀ ( NHMUK) “ INDIA: Kurseong Div. Lat Panchar. 4,000 ft. vi.1934. H.G.Champion. // G.C.Champion Coll. B.M.1927 409.” ; 1♀ ( MNHN) “ Environs de Kurseong R P. Bretaudeau // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” ; 1♀ ( MNHN) “ Sikkim Rhenok Eté 1894 Chasseurs Bretaudeau” ; 1♀ ( MNHN) “ Sikkim Lanpamum // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” ; 1♂ ( NHMUK) “4 // VII-21-1923 El. 5000 ft. // Tendong Sikkim // NHMUK015015957” ; 1♂ ( MNHN) “ Maria Basti British Bootang. // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” ; 1♂ ( MNHN) “ British Bootang Sakiȯu L. Durel // Anomala flavonotata , arr. G.J.Arrow det. // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” ; 2♂♂, 2♀♀ ( MFPC) “ BHUTAN: Monger, Thebong 27°17′N 90°20′E 19.VII.2009 H. 2273 m Viktor Siniaev leg.” GoogleMaps ; 1♂ ( MFPC) “ China-India border: Tawang 2500 m 05–10. JUL. 2013 Takeshi Maeda leg.” ; 1♂ ( ZMPC) “ CHINA: Tibet, Shigatse , Lower Yatung, N 27°15′22.38′′, E 89°01′10.17′′, 1832 m, 2023.VII.10, at light, Quan-Yu Ji leg.”

Description. Male. Body shape elongated ovoid, moderately convex.

Color . Generally brown to black, elytra sometimes lighter; shiny, with weak greenish luster, head tinged with bronze luster; elytra with a yellow, narrow to broad zigzag band extending from primary costa 5 to 1, sometimes partly interrupted; antennae reddish or yellowish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown.

Head. Greatest width/length of clypeus approximately 2.2; subtrapezoidal, lateral margins straight and weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface with very dense, large and irregular punctures. Frontoclypeal suture straight, distinct. Anterior half of frons with dense and large punctures, gradually sparser and smaller posteriad. Interocular distance/maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1–6. Inner margin of eye with a few moderately long setae.

Pronotum. Sides strongly convergent anteriad and weakly curved. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line indistinct and broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and small punctures. Lateral margin with several long setae.

Scutellum triangular, lateral margin slightly excurved; with moderately dense and minute punctures.

Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, irregular and large punctures, interstice II with a somewhat irregular secondary stria, all intervals bearing moderately dense and minute punctures, most sparse on subsutural interstice. Humeral protuberance moderately prominent, apical protuberance strongly prominent. Lateral carina weakly expanded from base to apical two fifths. Epipleuron with a row of sparse, short to moderately long setae from base to apical fifth.

Abdomen. Propygidium with moderately dense and transverse punctures. Pygidium with dense and transverse striation, gradually breaking up into sparse and very transverse punctures towards the smooth greatest tumidity; with some long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1–3 strongly carinate laterally; ventrites 2–5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 with very dense and irregular punctures; ventrites 2–5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin.

Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur/narrowest extremity of mesosternum approximately 6.0.

Legs. Protibia bidentate, both teeth distinct; apical tooth acute or rounded apically, extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw not convex, forming a notch internobasally; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional rounded basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae.

Male genitalia. See figs 14A–H and 15A–D.

Female. Interocular distance/maximum head width approximately 0.7; length of antennal club approximately 0.8 times as long as combined length of antennomeres 1–6. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, subapical tooth somewhat rounded apically; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, basal denticle of metatarsomere 5 absent; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider and shorter than upper branch.

Measurement. Body length: 14.7–15.3 mm in males and 14.9–15.4 mm in females, greatest width: 8.3–9.0 mm in males and 8.2–9.2 mm in females.

Differential diagnosis. Anomala flavonotata is characterized by a bifurcate upper branch of the left paramere, with a short side branch present in the apical third of the left paramere, combined with a simply pointed apex of the left paramere in lateral view. The internoapical angle is either absent or shortly produced in contrast to A. menyuensis sp. nov. Moreover, the epipleura are only sparsely covered with setae in A. flavonotata (densely covered in A. menyuensis sp. nov.).

Distribution. China ( Xizang) (new record); India ( West Bengal, Sikkim); Bhutan.