Anomala incurva Zhao, Fujioka & Zorn, 2025

Zhao, Ming-Zhi, Fujioka, Masayuki & Zorn, Carsten, 2025, Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae), Zootaxa 5693 (3), pp. 301-356 : 314-315

publication ID

https://doi.org/10.11646/zootaxa.5693.3.1

publication LSID

lsid:zoobank.org:pub:3F45C3AC-27B2-4423-A260-E6586E0E27BA

DOI

https://doi.org/10.5281/zenodo.17322489

persistent identifier

https://treatment.plazi.org/id/4A5987BA-FFAB-9313-E9CA-81A0FDEAFEDA

treatment provided by

Plazi

scientific name

Anomala incurva Zhao, Fujioka & Zorn
status

sp. nov.

Anomala incurva Zhao, Fujioka & Zorn View in CoL , sp. nov.

[Chinese vernacular name: 内卷异丽aek]

( Figs 4E View FIGURE 4 , 10F View FIGURE 10 , 18E–H View FIGURE 18 )

Type material. Holotype: ♂ ( NSMT) “ China-India border: alt. 2500m West Kameng 29.JUN. – 03. JUL. 2015 Takeshi Maeda-leg.”

Description. Male. Body shape elongated ovoid, moderately convex.

Color . Generally black; shiny, with weak bronze luster; elytra dark brown, with a yellow, broad to very broad zigzag band extending from primary costa 5 to 1; protibial teeth and antennae reddish brown; setae yellowish brown, robust setae reddish brown.

Head. Greatest width/length of clypeus approximately 2.3; subtrapezoidal, lateral margins almost straight and subparallel, anterior margin slightly convex, anterior corners obsolete and broadly rounded; anterior margin weakly reflexed; surface very densely and finely rugopunctate. Frontoclypeal suture straight, distinct. Anterior third of frons finely rugopunctate, other portions including vertex with moderately dense and large punctures, slightly sparser and smaller posteriad. Interocular distance/maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1–6. Inner margin of eye with a few moderately long setae.

Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third or posterior third, weakly curved at middle. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and large punctures. Lateral margin with several long setae.

Scutellum triangular, lateral margin moderately excurved; with dense and small punctures.

Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, moderately dense and large punctures, interstice II with a somewhat irregular secondary stria, all intervals bearing moderately dense and minute punctures. Humeral and apical protuberances both strongly prominent. Lateral carina weakly expanded from base to middle. Epipleuron with a row of rather sparse and short setae in basal third.

Abdomen. Propygidium with very dense and transverse striation. Pygidium with dense and transverse striation, gradually sparser apicad, breaking up into sparse and transverse punctures only near the greatest tumidity; with some long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1–3 strongly carinate laterally; ventrites 2–5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 very densely and finely rugostriolate; ventrites 2–5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin.

Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur/narrowest extremity of mesosternum approximately 6.0.

Legs. Protibia bidentate, both teeth distinct; apical tooth extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, both equal in length; lower margin of lower branch of inner protarsal claw not convex, forming a notch internobasally; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional rounded basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae.

Male genitalia. See fig. 18E–H.

Female. Unknown.

Measurement. Body length: 16.2 mm, greatest width: 8.2 mm.

Differential diagnosis. Anomala incurva sp. nov. is very similar to A. latifasciata sp. nov. But the upper branch of the left paramere bears a widely expanded, flat and incurved apex. The lower branches of both parameres have straight lateral margins.

Etymology. The species name derives from the Latin adjective incurvatus, meaning curved or bent, referring to the special shape of left paramere.

NSMT

National Science Museum (Natural History)

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Anomala

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