Anomala luoyuensis Zhao, Fujioka & Zorn, 2025

Anomala luoyuensis Zhao, Fujioka & Zorn View in CoL , sp. nov.

[Chinese vernacular name: Ḇā异丽aek]

( Figs 3C–F View FIGURE 3 , 10E View FIGURE 10 , 17A–H View FIGURE 17 )

Type material ( 20♂, 16♀♀). Holotype: ♂ ( SCAU) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill. , 1753 m, 29.214870°N, 95.169272°E, 18–30.VII.2023 (at light), Yao-Nan Zhang leg.” GoogleMaps Paratypes: 6♂♂, 8♀♀ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill. , 1753 m, 29.214870°N, 95.169272°E, 18–30.VII.2023 (at light), Yao-Nan Zhang leg.” GoogleMaps ; 5♂♂, 5♀♀ ( ZYPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill. , 1753 m, 29.214870°N, 95.169272°E, 18–30.VII.2023 (at light), Yao-Nan Zhang leg.” GoogleMaps ; 5♂♂, 2♀♀ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng, Gelin Vill. , 1768 m, 29.21315309°N, 95.17111894°E, 2023.VII.13-14, Hao-Yi Liu leg.” GoogleMaps ; 1♂ ( MFPC) “ China-India border: Upper Siang 1850m 04–10. AUG. 2013 Takeshi Maeda-leg.” ; 2♂♂, 1♀ ( MFPC) “ China-India border: alt. 1500m Dibang Valley 21–30. JUN. 2017 Takeshi Maeda-leg.”

Description. Male. Body shape elongated ovoid, moderately convex.

Color . Generally dark brown to black, the elytra usually lighter; shiny, with weak to strong greenish luster; elytra with a yellow, narrow to broad zigzag band extending from primary costa 5 to 1; antennae reddish or yellowish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown.

Head. Greatest width/length of clypeus approximately 2.3; subtrapezoidal, lateral margins straight and moderately convergent anteriad, anterior margin slightly convex, anterior corners obsolete and broadly rounded; anterior margin weakly reflexed; surface very densely and coarsely rugopunctate. Frontoclypeal suture slightly incurved, distinct. An inverted triangular area on anterior half of frons rugopunctate, other portions with dense and large punctures, slightly sparser and smaller posteriad. Interocular distance/maximum head width approximately 0.65. Length of antennal club slightly shorter than combined length of antennomeres 1–6. Inner margin of eye with a few moderately long setae.

Pronotum. Sides strongly convergent anteriad and weakly curved. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and large punctures. Lateral margin with several long setae.

Scutellum triangular, lateral margin moderately excurved; with dense and small punctures.

Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, irregular and large punctures, interstice II with a secondary stria, all intervals bearing moderately dense punctures, which are slightly smaller than strial ones. Humeral protuberance moderately prominent, apical protuberance strongly prominent. Lateral carina weakly expanded from base to apical two fifths. Epipleuron with a row of dense, moderately long setae from base to apical protuberance, often partly abraded.

Abdomen. Propygidium with very dense, transverse and very fine striation. Pygidium with dense and transverse striation, sparse only on the greatest tumidity; with some long setae preapically and a transverse row of long setae along posterior margin. Abdominal ventrites 1–3 strongly carinate laterally; ventrites 2–5 each with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated, denser laterad, frequently confluent at the lateralmost area, ventrite 6 very densely and finely rugostriolate; ventrites 2–5 with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin.

Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur/narrowest extremity of mesosternum approximately 6.0.

Legs. Protibia bidentate, both teeth distinct; apical tooth extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw not convex, forming a notch internobasally; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional triangular basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae.

Male genitalia. See fig. 17A–H.

Female. Length of antennal club approximately 0.8 times as long as combined length of antennomeres 1–6. Lateral margin of pronotum more excurved. Lateral carina of elytra slightly wider than in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, basal denticle of metatarsomere 5 absent; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw shorter than the upper one.

Measurement. Body length: 13.7–15.5 mm in males and 15.0– 16.2 mm in females, greatest width: 7.5–8.9 mm in males and 8.2–9.2 mm in females.

Differential diagnosis. The parameres of Anomala luoyuensis sp. nov. are most similar to those of A. spiniforceps sp. nov. in general structure. In A. luoyuensis , the upper branch of the left paramere is distinctly thickened and unbranched, and the lower branches of both parameres are completely narrowed in ventral view, rather long, unequal in length, and parallel. In contrast, the lower branches are broad in A. spiniforceps , with only the internoapical angle narrowed and protruding, equal in length and overlapping.

Etymology. Luoyu (also spelled Lhoyul; Ḇā in Chinese) is a regional name that has historically been used to refer to an area in southeastern Tibet, roughly corresponding to present-day Motuo County. The specific epithet refers to the distribution of the new species in this region.