Anomala dilatostilus Zhao, Fujioka & Zorn, 2025

Zhao, Ming-Zhi, Fujioka, Masayuki & Zorn, Carsten, 2025, Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae), Zootaxa 5693 (3), pp. 301-356 : 319-320

publication ID

https://doi.org/10.11646/zootaxa.5693.3.1

publication LSID

lsid:zoobank.org:pub:3F45C3AC-27B2-4423-A260-E6586E0E27BA

DOI

https://doi.org/10.5281/zenodo.17322497

persistent identifier

https://treatment.plazi.org/id/4A5987BA-FFB4-930E-E9CA-81A0FBAEFD57

treatment provided by

Plazi

scientific name

Anomala dilatostilus Zhao, Fujioka & Zorn
status

sp. nov.

Anomala dilatostilus Zhao, Fujioka & Zorn View in CoL , sp. nov.

[Chinese vernacular name: ẋĩ异丽aek]

( Figs 6A–C View FIGURE 6 , 11D View FIGURE 11 , 20A–D View FIGURE 20 )

Type material ( 11♂♂, 6♀♀). Holotype: ♂ ( SCAU) “ CHINA: Yunnan, Dehong, Yingjiang, Tongbiguan , 2019.IV– VI, at light Wei-Zong Yang leg.” Paratypes: 9♂♂, 6♀♀ ( ZMPC) “ CHINA: Yunnan, Dehong, Yingjiang, Tongbiguan , 2019.IV–VI, at light Wei-Zong Yang leg.” ; 1♂ ( CZPC) “ CHINA: Yunnan, Dehong, Yingjiang, Tongbiguan , 2019. IV–VI, at light Wei-Zong Yang leg.” .

Description. Male. Body shape elongated ovoid, moderately convex.

Color . Generally dark brown to black; shiny, with weak greenish luster; elytra with a yellow, narrow zigzag band extending from primary costa 5 to 1, usually broken up into three spots; antennae dark brown; protibial teeth reddish brown only at extremity; abdominal ventrites 2–4 sometimes with some vague reddish spots; setae yellowish brown, robust setae reddish brown.

Head. Greatest width/length of clypeus approximately 2.3; subtrapezoidal, lateral margins moderately convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface densely and coarsely rugopunctate. Frontoclypeal suture straight, distinct. An inverted triangular area on anterior third of frons with dense and coarse punctures, other portions with dense and large punctures, slightly sparser and smaller posteriad, sparse on vertex. Interocular distance/maximum head width approximately 0.6. Length of antennal club equally as long as combined length of antennomeres 1–6. Inner margin of eye with a few moderately long setae.

Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior and posterior third, curved at middle. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line indistinct and broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and large punctures. Lateral margin with several long setae.

Scutellum triangular, lateral margin weakly to moderately excurved; with dense and large punctures.

Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, irregular and large punctures, interstice II with a somewhat irregular secondary stria, all intervals with moderately dense punctures, which are slightly smaller than strial ones. Humeral protuberance moderately prominent, apical protberance strongly prominent. Lateral carina expanded from base to apical two fifths. Epipleuron with a row of moderately dense and short setae from base to near apex, often partly abraded.

Abdomen. Propygidium with very dense, transverse and fine striation. Pygidium with dense and transverse striation, sparse around the greatest tumidity; with some long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1–3 strongly carinate laterally; ventrites 2–5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated, denser laterad, frequently confluent at the lateralmost area, ventrite 6 moderately densely rugostriolate; ventrites 2–5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin.

Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur/narrowest extremity of mesosternum approximately 6.0.

Legs. Protibia bidentate, both teeth distinct; apical tooth extending to middle of protarsomere 2, preapical tooth rounded. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw convex distally next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional rounded basal protuberance. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae.

Male genitalia. See fig. 20A–D.

Female. The elytral band always formed by two or three separate small spots, sometimes very indistinct. Length of antennal club approximately 0.8 times as long as combined length of antennomeres 1–6. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, extending to middle of protarsomere 3, preapical tooth acute; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small and acute, basal protuberance of metatarsomere 5 absent; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw shorter than its upper one and the lower margin not convex.

Measurement. Body length: 14.7–16.0 mm in males and 15.3–15.5 mm in females, greatest width: 8.3–9.2 mm in males and 9.0 mm in females.

Differential diagnosis. Anomala dilatostilus sp. nov. can be distinguished from all other species of the A. flavonotata -subgroup by the longest upper branch of the left paramere, which is abruptly widened in the distal section. The lower branches of both parameres are long, with an almost straight inner margin.

Etymology. The name dilatostilus is derived from the Latin adjective dilatus (expanded or widened) and noun stilus (stem). It describes the abruptly widening of the left paramere in the distal section.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Anomala

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