Anomala flavofasciata Arrow, 1912

Anomala flavofasciata Arrow, 1912 View in CoL

[Chinese vernacular name: ƀŝ异丽aek]

( Figs 8D–F View FIGURE 8 , 12C View FIGURE 12 , 23A–D View FIGURE 23 )

Anomala flavofasciata Arrow, 1912: 72 View in CoL [original type locality: “Sikkim: Kurseong, 4700–5000 ft. ”, “Gantok”, “ Tonkin: Mauson Mts., 2000–3000 ft. ”]; Arrow 1917: 209; Zorn 2005: 306, figs 3–5 [ lectotype designation]; Zorn & Bezděk 2016: 333.

Type material. See Zorn (2005). Lectotype from Kurseong, paralectotype from Gantok.

Additional material examined ( 20♂♂, 31♀♀). 1♂ ( NMPC) “ NEPAL SE 28.6. - 1.7.2012 Shivalaya env. E. Kučera lgt. Nep/B/01” ; 1♀ ( NHMB) “ Phul Choki 2000 m 7.6. // Nepal, 1977 W Wittmer” ; 2♂♂, 1♀ ( NHMUK) “ NEPAL, 5800' Kathmandu Dist. Godawari, at light 5/vi/1983 // M.J.D.Brendell. BMNH {E} 1983-222.” ; 1♀ ( NHMB) “Kali-G. Khola Kopchepani 1600m 18.VI.1986 // W-Nepal Dhawalagiri C. Holzschuh ” ; 1♂ ( NHMB) “Kali-G. Khola Kopchepani-Gasa 1600–2000m 19.VI.1986 // W-Nepal Dhawalagiri C. Holzschuh ” ; 1♀ ( NHMB) “Kali-G. Khola Gasa-Kalopani 2000–2500m 20.VI.1986 // W-Nepal Dhawalagiri C. Holzschuh ” ; 1♂ ( NHMB) “Kali-G. Khola Kalopani 25/ 2800m 21–25.VI.1986 // W-Nepal Dhawalagiri C. Holzschuh ” ; 1♂ ( NHMB) “ NEPAL: Kosi - #13 Depitar 27°27′N / 87°17′E to Barabishe 27°26′N / 87°18′E 1250- 560m, 12.vi.01 // NHMB Basel expedition to Nepal, 2001” GoogleMaps ; 1♂ ( NMBF) “ NEPAL Likhu Khola Tal 1700 m 4.VI.62 leg. G. Ebert // Anomala flaviventris Arr. ” ; 1♀ ( NMBF) “ Nepal, Kathmandu Valley Godavari 16-1800m 1.VI.1967 Dierl-Forster-Schacht // Anomala flaviventris Arr. det. G. Frey, 1973” ; 1♀ ( NMBF) “ Nepal, Kathmandu Valley Godavari 16-1800m 5.VI.1967 Dierl-Forster-Schacht // Anomala flaviventris Arr. det. G. Frey, 1973” ; 1♀ ( CZPC) “NEPAL-HIMALAYA Annapurna Geb. - Lamjung H. lg. Schmidt 1994 // Chamje-Tal 10.6. 1500 -1700 m” ; 1♂, 4♀♀ ( ZMPC) “ China: Tibet, Shigatse, Nyalam co., Zhangmu , 1900 m, 2020.VII.29, Yu-Chen Zheng leg.” ; 5♀♀ ( ZMPC) “ China: Tibet, Shigatse, Nyalam co., Zhangmu , Yingbin Road 2000 m, 2020.VII.29, Yu-Chen Zheng leg.” ; 3♂♂, 3♀♀ ( ZMPC) “ China: Tibet, Shigatse, Nyalam co., Zhangmu Town , 1970 m 27.971656°N, 85.966972°E, light trap, 2023.VII.1 Zu-Qi Mai leg.” GoogleMaps ; 1♂ ( ZFMK) “ Pedong bei Darjeeling 1935 Sig. R. Oberthür Eing. Nr. 4, 1956. // Anomala flaviventris Arr. det. G. Frey, 1967/68” ; 1♂ ( MNHN) “ Pedong A. Desgodins // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” ; 1♂, 2♀♀ ( MNHN) “ Phedong R. P. Desgodins // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” ; 1♀ ( MNHN) “ British Bootang L. Durel 1898 // Anomala flavofasciata , a. Compared with type. G.J.A. // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” ; 1♂ ( MNHN) “ British Bootang Maria Basti 1899 // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR” ; 1♂ ( MNHN) “British Bootang Maria Basti L. Durel” ; 1♀ ( NMBF) “8272 18 // 1913 345 // Annandale Kurseong E. Himalayas alt. 4700-5000 ft. 28.VI-10. // Anomala flavofasciata Arrow ” ; 2♀♀ ( CZPC) “ INDIA – SIKKIM west PEELING env., 2100m 18.- 20.7.1997 Jan Schneider lgt.” ; 1♂ ( CZPC) “north INDIA- West Bengal Shingalila national park Shirikhola 18.- 28.5.1999 lgt. E. Kučera ( 2600 m.m)” ; 1♂, 2♀♀ ( CZPC) “north INDIA –west SIKKIM Khecheopari lake ( 1800 m.m) 04.- 10.6.1999 lgt. E. Kučera ” ; 1♀ ( CZPC) “ INDIA – SIKKIM Khecheopari Lake 21.6. – 26.6.2003 lgt. E. Kučera ” ; 2♀♀ ( MFPC) “ INDIA - SIKKIM Khecheopalri lake 15.6– 22.6.2008 1800 m leg. E. Kučera ” ; 1♀ ( NMBF) “ 21 km O Wangdi Phodr. [ Wangdue Phodrang] 1700–2000 // Nat.-Hist. Museum Basel - Bhutan Expedition 1972 // Anomala spiloptera det. G. Frey, 1973” ; 2♂♂ ( GSPC) “ 21 km O Wangdi Phodr. 1700–2000 // Nat.-Hist. Museum Basel - Bhutan Expedition 1972 // Anomala spiloptera det. G. Frey, 1973” .

Description. Male. Body shape elongated ovoid, somewhat subparallel, moderately convex.

Color . Generally yellowish brown, distal and proximal sections of pro- and mesotibiae darker, the head, disc of pronotum except for a narrow lateral area, meso- and metathoracic surface, pygidium in anterior two thirds, abdominal ventrites 2–5 (in posterior half), metatibia and all tarsomeres reddish brown; dull, with weak iridescent sheen; elytra lighter than pronotum, with a yellowish brown, narrow to broad zigzag band extending from primary costa 4 to 1, sometimes very pale or partly interrupted; setae yellowish brown, robust setae reddish brown.

Head. Greatest width/length of clypeus approximately 2.1; subtrapezoidal, lateral margins weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed;

surface with dense, large and coarse punctures. Frontoclypeal suture slightly incurved, distinct. An inverted triangular area on anterior half of frons punctate as clypeus, other portions with very dense and large punctures, smaller posteriad. Interocular distance/maximum head width approximately 0.65. Length of antennal club 0.8 times the combined length of antennomeres 1–6. Inner margin of eye with a few moderately long setae.

Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third and posterior third, moderately excurved at middle. Anterior angle acute and protruding, posterior angle obtuse. Posterior marginal line absent, other marginal lines complete. Lateral fovea indistinct. Disc with dense, rather large and transverse punctures, smaller and rounded along lateral and posterior margins. Lateral margin with several long setae.

Scutellum triangular, lateral margin moderately excurved, or semielliptical; with dense and large punctures as well as several erect and moderately long setae.

Elytra. Surface weakly wrinkled, intervals moderately elevated, subsutural interstice widest; strial punctures dense and large, annulated, subsutural interstice with irregularly doubled secondary stria, interstices II–IV each with a somewhat irregular secondary stria, all intervals with moderately dense punctures, which are slightly smaller than strial ones, each bearing a minute seta. Humeral and apical protuberance moderately prominent. Lateral carina slightly expanded from base to apical third. Epipleuron with a row of moderately dense and moderately long setae from base to apical two fifths, usually partly abraded.

Abdomen. Propygidium with dense, transverse and fine striation. Pygidium with moderately dense and transverse striation, sparse only on the greatest tumidity; with sparse and long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites not carinate laterally; ventrites 2–5 with transverse and incompletely annulated punctures, short and moderately dense medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 densely rugostriolate; ventrites 2–5 each with a transverse and moderately dense row of long setae, dense laterally, the row of ventrite 6 present at posterior margin.

Ventral thoracic surface. Hypomeron with sparse and small punctures, moderately dense and long setae. Ventral mesothoracic surface with dense and small punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and long setae, medial area with smaller punctures, density and length of setae as lateral area. Metacoxa punctate as lateral metasternum but bearing sparser setae. Very narrow at the extremity of mesosternum.

Legs. Protibia bidentate, both teeth distinct and acute; apical tooth extending to middle of protarsomere 2. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches equally as long as and two times wider than their upper branches; lower margin of lower branch of inner protarsal claw not convex next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 smaller. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and rather long setae between rows; ventral surface of metafemur with two rows of moderately dense and long setae, one lateral and the other submedial, also with some irregular and long setae sublaterally; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae.

Male genitalia. See fig. 23A–D.

Female. Generally reddish or blackish brown, lateral area of abdominal ventrites, mesofemur, marginal areas of metafemur, as well as antenna yellowish brown, protibial teeth reddish brown; elytra not paler than pronotum, zigzag band narrower, sometimes breaking up into three small spots.

Antenna shorter, length of antennal club approximately 0.8 times the combined length of antennomeres 1–6. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, extending to anterior margin of protarsomere 2; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of protarsomere 5 distinctly smaller than in male; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch.

Measurement. Body length: 12.6–14.2 mm in males and 12.9–14.4 mm in females, greatest width: 7.1–7.8 mm in males and 6.9–8.2 mm in females.

Differential diagnosis. The ventral surface of the parameres is gradually changing from sclerotized to membranous in the apical half in Anomala flavofasciata . Compared to similar species, the parameres are also most slender.

Remarks. The record of A. spiloptera Burmeister, 1855 from Wangdi Phodrong, Bhutan, published by Frey (1975), was shown to be A. flavofasciata after examination of the original specimens (NMBF and GSPC, with Frey’s label).

Distribution. China ( Xizang) (new record); India ( West Bengal, Sikkim); Nepal; Bhutan.