Cymatodera insalebrosa Rifkind, 2020

Cymatodera insalebrosa Rifkind , new species

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( Figs. 4 View Figs , 35, 36 View Figs )

Specimens Examined. Holotype male: Mexico, Puebla, Zapotitlan, VI-28-1965, Burke, Meyer, Schaffner ( TAMU) . Paratypes: 1 M ( CNIN), 1 F ( TAMU), same data as holotype ; MEXICO: PUEBLA: 1 M ( JNRC), 6 mi SW Tehuacan , July 7, 1973, taken at light, Mastro & Shaffner ; OAXACA: 1 M ( EMEC), 3 mi SW El Tule , IV-12-1958, R. C. Bechtel, E. I. Schlinger, collectors .

Description ( Holotype). Length: 13.0 mm. Color: Elytral ground color brown; head and pronotum darker, reddish brown; mouthparts, antennae, and venter reddish testaceous; elytra with three sets of eburneous markings ( Fig. 4 View Figs ), only the middle pair attaining the lateral margins. Head: Eye moderate in size; antenna elongate; antennomeres subserrate: Antennomere 2 about 3/4 the length of antennomere 3; antennomere 11 about as long as antennomere 10, rounded at apex; surface rather closely, finely but shallowly punctate, densely but inconspicuously set with fine, very short, reclinate pale setae, intermingled with fewer, longer, more robust, erect pale setae. Pronotum: Elongate (ratio of length to width 13:9); subflattened on disc above; moderately densely but shallowly punctate, transversely rugulose, rather sparsely set with fine, suberect pale setae of moderate length, interspersed with erect, more elongate and more robust, testaceous setae. Elytra: Elongate (together more than 2× as long as broad), umbones well developed, sides subparallel, gradually, arcuately convergent from posterior 1/4 to slightly dehiscent apices; surface shining, very shallowly roughened, strial punctures rather minute, darkened, not set in close proximity to one another; vestiture rather sparse, composed mostly of rather short, suberect, pale testaceous setae, intermixed with fewer longer, more robust erect setae. Metaventrite: Shining, minutely shallowly punctulate; carinae absent; tubercles prominent, closely set on either side of midline posteriorly; surface with a dense covering of fine, tawny setae of moderate length. Abdomen: Ventrites 1–5 with a large, shallow, obovate depression on each side, surface shallowly dimpled and inconspicuously punctulate, moderately densely but inconspicuously clothed with fine, suberect pale tawny setae; ventrite 5 ( Fig. 35 View Figs ) with hind angles obtuse, posterior margin sinuate laterally, deeply, arcuately emarginate at middle; ventrite 6 ( Fig. 35 View Figs ) much longer than broad, sides slightly convergent posteriorly, ventrolateral margins carinate, extending to and circumscribing and defining rounded (non-produced) posterior angles and rather broad, subtruncate posterior margin; middle of posterior margin broadly triangularly emarginate; ventral surface of ventrite 6 shallowly roughened, raised at middle where it bears a distinct longitudinal carina that extends from anterior to posterior margins; tergite 5 with posterior margin rather acutely inflected laterally, arcuately emarginate at middle; tergite 6 ( Fig. 36 View Figs ) narrower and slightly shorter than sternite 6, sides broadly arcuate, slightly convergent posteriorly, surface granulate, with a shallow longitudinal carina at middle, hind margin shallowly, sub-triangularly emarginate. Aedeagus: Apex of phallus upturned, sagittate.

Variation. Specimens range in length from 10–13 mm. The available female specimen has abdominal ventrite 5 with the sides oblique and the posterior margin with a broad U-shaped emargination at middle; tergite 6 and ventrite 6 are both rounded posteriorly, with tergite 6 slightly surpassing ventrite 6 and its margin very feebly inflected and depressed at middle.

Etymology. The specific name means “unroughened” and refers to the smooth integument of the metaventrite in this species.

Distribution. The new species is known from the Mexican states of Puebla and Oaxaca.

Biology. One specimen was collected at light.

Diagnosis. Based on size, general facies, presence of dense metaventral setation (males), and the shape of the pygidia, Cymatodera insalebrosa appears most similar to Cymatodera bellamyi Rifkind and Cymatodera cicatricula Rifkind , both also known from southern Mexico, and potentially sympatric with the new species. Cymatodera insalebrosa , though possessing a dorsal elytral pattern similar to C. bellamyi , has much finer elytral punctation. Males of C. insalebrosa may be easily separated from these congeners by the sculpturing of their metaventral integument, which is smooth and bears a distinct pair of posterior tubercles. By contrast, the metaventrite is asperate and lacking tubercles in the other species.