Josa chazaliae ( Simon, 1897 )

Josa chazaliae ( Simon, 1897) View in CoL

Figures 1–8 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4 View FIGURE 5 View FIGURE 6 View FIGURE 7 View FIGURE 8

Tomopisthes chazaliae Simon, 1897: 11 (female lectotype MNHP 18396 View Materials ; vchLAM-243 and two females paralectotypes MNHP vchLAM-242 and vchLAM-244 here designated, from Sierra Nevada de Santa Marta , Magdalena, Colombia, De Dalmas coll., in MNHP, examined).

Josa chazaliae View in CoL : Ramírez, 2003: 109.

Note. The syntypes correspond to two different species; the lectotype ( MNHP 18396 vchLAM-243) and one of the paralectotypes belong to Josa chazaliae ( MNHP vchLAM-242) and the other to an undescribed species, which is described below as Josa samaria sp. nov. ( MNHP vchLAM-244). Only records from the additional material examined were mapped, as the type’s label lacks a specific locality within the Sierra Nevada de Santa Marta.

Other material examined: COLOMBIA. Magdalena: Ciénaga, San Pedro, Parque Natural Nacional Sierra Nevada de Santa Marta , Loma de la Cebolleta , misc. vegetation, low-medium, manual caught, 3505 m, J. Kochalka leg., 9.V.1975, 1 ♂ (MUSENUV-Ar-2671; vchLAM-245), 1 ♂ (MACN-Ar 46690; vchLAM-246) . Cesar: Valledupar, Parque Nacional Natural Sierra Nevada de Santa Marta ( NW Donachuí), Meoyaca , Río Donachuí , 3974 m, [10°47′39.9″N, 73°36′03.6″W], 18.I.1974, 1 ♀ (MUSENUV-Ar-2672; vchLAM-247); GoogleMaps 3596 m, [10°47′04.5″N, 73°33′40.4″W], 2.I.1973, 2 ♀ (MACN-Ar 46691; vchLAM-248); GoogleMaps Donachuí, Rio Donachuí , top of woody vegetation, 4230 m, [10°47′50.0″N, 73°37′34.5″W], 6.I.1973, 1 ♀ (MUSENUV-Ar-2673) GoogleMaps .

Diagnosis. Males of Josa chazaliae are similar to those of Josa berlandi sp. nov. by the small conductor with the external region apically bifid and by the short embolus ( Figs 7H–I View FIGURE 7 , compare with Figs 25A–D View FIGURE 25 , 26A–C View FIGURE 26 , 27A–I View FIGURE 27 ), but can be differentiated from this species by the conductor widening towards the apex with both apical tips of the external section of the conductor of about the same length, the posterior one curved towards the internal region and with a subapical pronounced bump ( Fig. 7A–D View FIGURE 7 , arrow), median apophysis bifid with its anterior branch tubular and slightly curved ( Figs 5A View FIGURE 5 , 6A View FIGURE 6 , 7A–D View FIGURE 7 ), and the very small and rugous retrobasal tegular lobe ( Fig. 7A, F View FIGURE 7 ). Females are easily distinguishable by the very thin and short ducts of the accessory bulb restricted to the lateral regions of the vulva ( Figs 6D–E View FIGURE 6 , 8A–D View FIGURE 8 ), the swollen, triangular basal sclerotized projections of the epigyne ( Figs 2E View FIGURE 2 , 5E View FIGURE 5 ), the long median septum, and median field oval with deep copulatory openings ( Figs 2E View FIGURE 2 , 5E–F View FIGURE 5 , 6D–E View FIGURE 6 , 8A–F View FIGURE 8 ).

Description. Female (description based on the lectotype, MNHP 18396; vchLAM-243 and a female MUSENUV- Ar-2671; vchLAM-245). ( Figs 1–2 View FIGURE 1 View FIGURE 2 , 4 View FIGURE 4 ): Total length 7.42. Carapace length 3.70, width 2.56, height 1.64. Clypeus length 0.22. Sternum length 1.88, width 1.29. Legs. Leg I: ti length 1.90/width 0.51, mt 1.53. II: ti 1.81/0.47, mt 1.49. III: ti 1.61/0.41, mt 1.57. IV: ti 2.21/0.46, mt 2.18. Spines: legs I–II, femur d 1-1-1, p 2ap; tibia v 2-2-2; metatarsus v 2bas. III, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-0-2, p d1-1-0-1, r d1-1-0-1. IV=III but femur r d1ap; patella d 1-1, r 1; metatarsus v 2-2-2. Abdomen: length 3.75, width 1.86, spiracle–epigastrium 1.26. Body dehydrated, damaged. Color: carapace dark brown, with two wide longitudinal dark bands on sides, narrowed in cephalic region. Ocular area on dark patch with eyes bordered by black rings. Carapace with fringe of white setae on sides. Clypeus brown, low, with some white setae. Chelicerae brown with black patches and fine setae on front. Endites, labium, and sternum dark brown, labium and endites light in apical region. Legs dark brown, with long darker stripes on dorsal edges of femora. All articles covered by white setae. Abdomen dorsally with median spots on dark gray background and abundant white setae; anterior region with bunch of thick setae; venter uniformly dark gray, with light dots. Epigyne: anterior plate sclerotized with deep anterior excavations, median notch long and narrow ( Figs 2E View FIGURE 2 , 5E View FIGURE 5 , arrows). Median field transverse, large, oval in posterior view, with medial depression ( Fig. 8E–F View FIGURE 8 ). Lateral lobes with sclerotized extensions very wide, forming a longitudinal concavity, posterior extensions rounded at base ( Figs 2E View FIGURE 2 , 5E View FIGURE 5 ). Copulatory ducts long, touching with the accessory bulb ducts medially ( Fig. 6E View FIGURE 6 ). Spermathecae rounded medially positioned in the epigyne, oriented towards median region ( Figs 5F View FIGURE 5 , 6E View FIGURE 6 , 8A–E View FIGURE 8 ). Ducts of accessory bulbs long, thin, convergent ( Figs 2E View FIGURE 2 , 5E–F View FIGURE 5 , 6D–E View FIGURE 6 , 7 View FIGURE 7 ).

Male (MUSENUV-Ar-2671; vchLAM-245) ( Figs 3 View FIGURE 3 ): Total length 4.54. Carapace length 2.34, width 1.61, height 0.72. Clypeus length 0.10. Sternum length 1.20, width 0.79. Legs. Leg I: ti length 2.17/width 0.26, mt 1.88. II: ti 1.97/0.25, mt 1.72. III: ti 1.32/0.23, mt 1.20. IV: ti 1.62/0.27, mt 1.55. Spines: legs s I–II, femur d 1-1-1, p 2ap, r 0-d1-d1; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0; metatarsus v 2-2-0, p d1-d1-0, r d1-d1. III–IV, femur d 1-1-1, p 0- d1-d1, r 0-d1-d1; patella r 1me; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-2-2, p d1-1-0-1, r d1-1-0-1. Abdomen: length 2.34, width 1.35, spiracle–epigastrium 0.83. Color: carapace brown, with two faint dark bands on sides, anteriorly covered by white setae. Ocular area brown with eyes bordered in black rings. Clypeus brown, low, with abundant white setae. Chelicerae brown with black stripes and fine setae on front. Endites, labium, and sternum brown; labium and sternum darker. Legs coxae-tarsi brown, with some tenuous dark patches. Articles with a few white and black setae. Abdomen dehydrated, with a pattern of faint spots on a brown background. Dorsum covered with few fine white setae. Venter uniformly brown. Palp: tegulum with small basal retrolateral lobe, very sclerotized, rugous ( Figs 5A View FIGURE 5 , 6A View FIGURE 6 , 7A View FIGURE 7 , 8A–F View FIGURE 8 ). Median apophysis large, bifid, anterior tip long, tubular, hook-shaped, posterior branch membranous at base with external border sub-rounded ( Figs 5A View FIGURE 5 , 6A View FIGURE 6 , 7A–B View FIGURE 7 ). Conductor small, rounded, with projected base, external section wide, anteriorly swollen, bifid, with both tips equal in length, but posterior tip curved towards internal region ( Fig. 7A–B, D View FIGURE 7 ); internal section ending as a sharp tip ( Fig. 7D View FIGURE 7 ). Paramedian apophysis cup-shaped, small, heavily sclerotized, concave, apically sharp ( Figs 5A, D View FIGURE 5 ; 6A–C View FIGURE 6 ; 7A–E View FIGURE 7 ). Embolus short, not describing a complete loop around bulb, tape-shaped at base, coiled, with poorly developed laminar projection; basal process small, flat, retrolateral region pointed in posterior region, prolateral region oval, poorly pronounced into external keel ( Figs 5A–D View FIGURE 5 , 6A–C View FIGURE 6 , 7 View FIGURE 7 ).

Variation: Males (n=2): total length: 4.28–4.54; carapace length: 2.18–2.34, width: 1.57–1.61; abdomen length: 2.14–2.34, width: 1.19–1.35. Females (n=3): total length: 7.40–8.19; carapace length: 3.52–3.70, width: 2.47–2.56; abdomen length: 3.75–4.42, width: 1.86–2.64. Fresh specimens look more pigmented.

Distribution. Only known for Sierra Nevada de Santa Marta, Colombia ( Fig. 31 View FIGURE 31 ).