Aikahika veta, Sroka & Agnihotri & Singh, 2025

Aikahika veta sp.nov.

( Figs 1, 2)

Zoobank registration: htp://zoobank.org/ urn:lsid:zoobank.org:act:

Diagnosis (male imago): Small size (body length c. 2.5 mm); penis lobes narrowing apically, separated from the base (distance between penis lobes slightly smaller than penis lobe width); large compound eyes; ratio fore tibia length/fore tarsus length c. 1.22; forceps widened subapically, with minute apical segment(s).

Etymology: Veta (Latin) means ‘old’, referring to the ancient age of the fossil.

Holotype: Accession number BSIP 42401.

Description

Body length 2.44 mm. Ventrolateral aspect observable from one side.

Head: Large compound eyes (diameter c. 0.39 mm), upper portion with rectangular facets ( Fig. 1A, B). Antennae filiform, 0.73 mm long, enlarged scapus slightly shorter than pedicellus.

Torax : Toracic sterna partially visible from ventrolateral view, with mesothoracic basisternum and furcasternum recognizable ( Fig. 1A). Furcasternum with median longitudinal invagination. Visibility of exact shape of sternites impaired by crack in amber matrix, creating optically distorting interface in this area. Right foreleg atached to body ( Fig. 1A, D), all segments complete except for tarsus; distal part of tarsus missing ( Fig. 1F). Length of fore femur 0.63 mm, fore tibia 0.94 mm long (ratio fore tibia/ fore femur length 1.54). Lef foreleg mostly missing ( Fig. 1A, D), isolated tarsus preserved with distal portion of tibia ( Fig. 1F). Length of tarsus 0.77 mm (ratio fore tibia/fore tarsus length 1.22). Tarsus five-segmented; segment 1 shortest, 0.05 mm long; segments 2 and 3 almost equal in length, 0.23 mm and 0.22 mm long, respectively; segment 4 is 0.17 mm long; and segment 5 is 0.1 mm long. Double tarsal claws indistinctly recognizable. Right middle leg complete, detached from body and positioned close to right foreleg ( Fig. 1A, D). Distal part of femur not visible, as well as proximal parts of tibia and tarsus. From lef middle leg, coxa and trochanter present ( Fig. 1A, D). Distal part of femur is visible close to body, tibia and tarsus missing. Right hind leg mostly missing, only coxa, trochanter, and proximal fragment of femur present, atached to body ( Fig. 1A, D). Lef hind leg complete, tibia fractured and deformed at places ( Fig. 1A, C–E). Hind femur 0.75 mm long, hind tibia 0.97 mm long (ratio hind tibia/hind femur length 1.29). Tibiopatellar suture not distinguishable. Tarsus four-segmented, 0.24 mm long (ratio hind tibia/hind tarsus length 4.04); segments 1–3 of similar length, segment 4 shortest. Double tarsal claws; one claw obtuse ( Fig. 1E). From wings, only lef forewing is preserved and partially visible, 2.46 mm long ( Figs 1A, 2A, B). Veins C, ScP, and R simple, well-observable, except distal portion. Costal brace ScA visible. RP bifurcated 0.63 mm from wing base. Five longitudinal veins in RP sector. Two short sections of main longitudinal veins visible posteriorly from RP, representing basal portions of MA and MP. Distal branching of median veins, cubital and anal veins not observable. Sparse crossveins present between longitudinal veins. Hindwings obscured, not observable.

Abdomen: Elongated, bent upwards, partially hidden by crack in matrix ( Fig. 1A). Genitalia well visible ( Fig. 1G, H). Two elongated penis lobes, parallel, separated from base. Distance between penis lobes slightly narrower than penis lobe width at base. Penis lobes narrowing apically, slightly bifurcated at apex. Forceps ( Fig. 1G, H) curved inwards, borders between segments not clearly distinguishable. Basal thicker portion of forceps either composed of two segments, approximately equal in length, or consisting of single segment constricted in middle. Apical segment short and narrow (possibility of two very short apical segments not excluded). Subgenital plate entire, without medial notch ( Fig. 1G, H). Two caudal filaments present, representing paracercus and one of cerci. Second cercus missing; point of its insertion visible at tip of abdomen ( Fig. 1I). Cercus 4.42 mm long (1.8× longer than body) and paracercus 2.29 mm long (approximately equal to body length). Paracercus relatively thick terminally and most probably incomplete. Both filaments covered by short, hair-like setae.

Atalophleboculata sp. (specimen BSIP 42402)

( Fig. 3)

Description Body length 3.17 mm. Lateral aspect observable from both sides.

Head: Large compound eyes (diameter c. 0.56 mm), upper portion with rectangular facets ( Fig. 3B, D). Antennae filiform, scapus and pedicellus not visible, flagellum c. 7.2 mm long.

Torax : Toracic sclerites not recognizable. Right foreleg mostly missing, only basal part preserved. Lef foreleg completely preserved ( Fig. 3A, B, J). Femur 0.65 mm long, tibia 1.28 mm long (ratio tibia/femur length 1.97). Length of tarsus 1.25 mm (ratio tibia/tarsus length 1.02). Tarsus five-segmented ( Fig. 3J); segment 1 shortest, 0.09 mm long; segments 2 and 3 of similar length, 0.49 mm and 0.35 mm long, respectively; segment 4 is 0.19 mm long; and segment 5 is 0.13 mm long. Double tarsal claws, one obtuse. Middle and hind legs completely preserved ( Fig. 3A–C, K, L). Middle leg with femur 0.65 mm long, tibia 0.57 mm long, and tarsus 0.27 mm long (ratio tibia/femur length 0.88, ratio tibia/tarsus length 2.11). Hind leg with femur 0.75 mm long, tibia 1.08 mm long, and tarsus 0.25 mm long (ratio tibia/femur length 1.44, ratio tibia/tarsus length 4.32). Individual tarsal segments only partially distinguishable in middle and hind leg ( Fig. 3K, L). Double tarsal claws (one obtuse) in middle and hind leg. Forewing c. 3.2 mm long, 1.12 mm wide ( Fig. 3A–C, E–G). Costal brace ScA present. Veins C, ScP, and R simple. Pterostigma with six thick crossveins observable in lef forewing. RP bifurcated c. 1.05 mm from wing base. Four longitudinal veins in RP sector. MA bifurcated more distally than RP, c. 1.32 mm from wing base; single intercalary vein between MA branches. Two MP branches, MP1 and MP2; possibly MP bifurcation basally or MP branches independent from base ( Fig. 3G). Single intercalary vein between MP branches. CuA recognizable as simple convex vein posteriorly from MP2. Sparse crossveins present between longitudinal veins. Hind wing more clearly visible on lef side ( Fig. 3H, I), minute, 0.3 mm long, with pronounced triangular pointed costal projection, 0.12 mm high. Venation faintly recognizable, consisting of simple subcostal, branched radial, and simple medial veins.

Abdomen: Narrow, elongated. Genitalia faintly visible from lateral view ( Fig. 3B). Forceps curved inwards. Tree caudal filaments (cerci and paracercus) of approximately same length, 2.36–2.44× longer than body, covered by short hair-like setae ( Fig. 1A).