Euglyptus punctatus (Broun, 1893)

Euglyptus punctatus (Broun, 1893) View in CoL

( Figs 12 View Fig , 16 View Fig )

Microtyrus punctatus Broun, 1893a:1413 View in CoL (#2461). Type locality: Maketu, Hunua Range [AK].

Microtyrus punctatus View in CoL : HUTTON (1904: 176); HUDsON (1923: 364); KUsCHEL (1990: 28); NEWTON & CHANDLER (1989: 21).

Euglyptus punctatus View in CoL : RAFFRAY (1904:550), RAFFRAY (1908:52), RAFFRAY (1911: 14); NOMURA & LEsCHEN (2006: 247).

Euplectus iracundus Broun, 1893a: 1429 View in CoL (#2481). Type locality:Maketu, Hunua Range [AK]; syn. nov.

Euplectus iracundus View in CoL : RAFFRAY (1908: 88), RAFFRAY (1911: 35); HUTTON (1904: 180); HUDsON (1923: 366).

Microtyrus iracundus : KUsCHEL (1990: 28).

Euglyptus iracundus View in CoL : NOMURA & LEsCHEN (2006: 247).

Type material. Microtyrus punctatus . LECTOTYPE (by present designation): “[red-bordered disc] Type / 1977 / Hunua (Maketu) / New Zealand [with red underline] Broun Coll. Brit. Mus. 1922-482. / Microtyrus punctatus .” ( BMNH, NHMUK015009957 ).

Euplectus iracundus . HOLOTYPE: ♀, “[red-bordered disc] Type / Hunua Maketu / New Zealand [with red underline] Broun Coll. Brit. Mus. 1922- 482. / Microtyrus iracundus [sic]” ( BMNH).

Additional material examined. NEW ZEALAND: NORTH ISLAND: AK [ Auckland]: Mt. William Scenie Reserve, Kauri Ridge Track, 3.2 km N Pokeno 94m, III-29-2010 D.S Chandler, sift kauri podocarp, & etc. forest litter // Compared with type of Broun Euplectus iracundus ♀ by DSChandler [green label] (7 JJ 4 ♀♀; DSC). Waitakere Ranges Reg. Pk., Upper Huia Reser. Track, II-28-2010, 4.3 km SW Waiatarua 277 m, D.S. Chandler, sift forest litter (5 JJ 1 ♀; DSC / 1 J; NZAC). Ex.Broun Collection. [handwritten] // Hunua Rg., Maketu.Auckland. // 2481. // Euglyptus iracundus Broun. [handwritten] // T.Broun Collection // A.E.Brookes Collection ( 2 ♀♀; NZAC). 2461 [handwritten] // T.Broun Collection // A.E.Brookes Collection // Possible Broun SYNTYPE Microtyrus punctatus D S Chandler 2010 [green label] (2 JJ; NZAC). BP [ Bay of Plenty]: Papatea 18 Oct 1992 J.S. Dugdale // Sifted litter 92/66 ( 2 ♀♀; NZAC). Papatea 19 Oct 1992 J.S. Dugdale // Sifted litter 92/66 (1 J slide mounted, 2 JJ genitalia dissected; NZAC). Papatea 15 Sep 1992 G. Hall & R.C. Henderson // Sifted litter 92/48 (1 J genitalia dissected, 1 ♀; NZAC). Papatea 5 Nov 1993 R.C. Henderson Litter 93/116 (1 J; NZAC). Papatea 19-22 Sep 1992 J.S. Dugdale Malaise trap (1 J; NZAC). Mt.Te Aroha, 3 km E Te Aroha, Dog Kennel Flat Track, 850 m, II-15-2010, D.S Chandler (4 JJ 1 ♀; DSC). Lottin Pt Rd Waenga Bush 16 Sep 1992 G.M.Barker (1 J; NZAC). CL [Coromandel]: Mt Moehau 7 Nov 1978 B.M.May moss 78/276 (1 J slide mounted; NZAC). RI [Rangitikei]: Pohangina Valley Totara Reserve 24 Dec 1980 J.C.Watt // Litter 80/164 ( 1 ♀; NZAC). TK [ Taranaki]: Mt Messenger 610’ Taranaki 24.xi.68.J.S.Dugdale ( 1 ♀; NZAC). WA [Wairarapa]: Castlehill / Haunu area, SE Puketoi Ra March 2010 (1 J genitalia dissected; JTN). WN [ Wellington]: Orongorongo Res.Stn Well. // 21May69 J.S.Dugdale (1 J genitalia dissected; NZAC). Pakuratahi Forks Kaitoke 27-Apr-97 // In forest leaf litter (1 J; JTN).Pakuratahi Forks Kaitoke, 1.8.93 // In decayed wood (1 J genitalia dissected; JTN). Greens Road Tiritea 2-Jun-04 // In rewarewa litter on stream terrace (1 J; JTN). Rimutuka Summit Tk, 15 Apr 2005, R. Leschen, C. McGuiness, // leaf litter, Nothofagus, Mahoe, RL 974, 41.06S, 175.13E (1 J; NZAC).Tinakori Hill Wellington, 5.9.94 // In bole of dead fivefinger // 614 (1 J genitalia dissected; NZAC). nikau Res Paraparaumu 23.4.88 (1 J genitalia dissected; JTN). Wilton Bush 25.5.92 (1 J genitalia dissected; JTN).Kaitoke 17.7.88 (1 J; JTN).Wilton Bush Wellington 3.7.88 (1 J genitalia dissected; JTN). WO [ Waikato]: Te Kauri Scenic Res. Nr Kawhia 20 Sep 1981 C.F.Butcher // litter 81/89 ( 1 ♀; NZAC). Herangi Ra, 500m, above Mangatoa Saddle 9 Dec 1982 // J.S. Dugdale Litter 82/136 (1 J genitalia dissected; NZAC). SOUTH ISLAND: NN [ Nelson]: Upper Maitai 13.2.57 E.S.Gourlay (1 J genitalia dissected, 1 J; NZAC). Upper Maitai, 3.V.1950 E.S. Gourlay leg. (1 J genitalia dissected; NZAC). Slaters Road, 0.7km s Whangamoa Saddle, 410m 13.xii.1984 - 4.i.1985 Nothofagus spp forest A.Newton/M.Thayer 703 // flight intercept (window) trap (1 J genitalia dissected; FMNH). SD [ Marlborough Sounds]: Tennyson Inlet, east side Duncan Bay, 30m 15.xii.1984 - 5.i.1985 podo.- Nothofagus for. A.Newton/M.Thayer 709 // flight intercept (window) trap (3 JJ genitalia dissected; FMNH). Tennyson Inlet, west side Te Mako Bay, 125m 15.xii.1984 - 5.i.1985 Nothofagus-podo-hdwd.A.Newton/M.Thayer 710 // flight intercept (window) trap (1 J; DSC / 1 J genitalia dissected; FMNH). D’Urville I Kapowai Apr 1971 F.Alack // Litter 71/101 (1 J; NZAC). Unknown: 31.viii.51. moa woodholm ni-. N.I. P182 [handwritten, illegible] (1 J; NZAC).

Diagnosis. Males of this species can be distinguished from other Euglyptus species by the following combination of characters: body 1.16–1.26 mm (forebody 0.94–0.98 mm) ( Fig. 12A View Fig ); head punctate ( Fig. 12A View Fig ); antennomere X as long as wide, subhexagonal, XI wider than X ( Fig. 12C View Fig ); pronotum gradually narrowing from middle to posterior 1/3, slightly narrowing from posterior 1/3 to posterior margin, with broad, shallow median longitudinal sulcus and broad transverse antebasal sulcus, with median antebasal carina extending from median antebasal impression to posterior margin ( Fig. 12B View Fig ); both parameres similar in length, angled ventrally, left paramere elongate and narrow, apex angled to right in ventral view, right paramere gradually narrowing to pointed apex, curved ventrally in lateral view; median lobe distinct, slightly sinuate, longer than parameres, evenly curving dorsally in lateral view, apical margin thin and fan-shaped ( Figs 12E–J View Fig ).

Redescription. Body 1.16–1.26 mm (forebody 0.94–0.98 mm); head, thorax, abdomen, and antennal scape reddish- -brown; elytra, leg, pedicel, and antennomeres III–VIII lighter; tarsus, maxillary palpus, and antennomeres IX–XI yellowish-brown ( Fig. 12A View Fig ).

Head wider than long, with small dorsal tentorial pits, impression weakly defined around dorsal tentorial pits; temples gradually narrowing toward posterior margin of head capsule, posterolaterally prominently angulate, with several setae; impression between antennal tubercles as wide as antennal tubercle diameter. Pedicel wider and longer than antennomeres III–VIII, antennomeres III–VIII as long as wide, subquadrate, VI and VIII smaller than others, IX–X as long as wide, subhexagonal ( Figs 12A, C View Fig ).

Pronotum as wide as head, slightly wider than long, narrowest at apex, gradually widening from anterior margin to middle, widest in middle, gradually narrowing from middle to posterior 1/3, slightly narrowing from posterior 1/3 to posterior margin, with broad, shallow median longitudinal sulcus and broad transverse antebasal sulcus, with median antebasal carina extending from median antebasal impression to posterior margin ( Fig. 12B View Fig ).

Elytra and abdominal tergites without any distinguishing characters ( Figs 12A, D View Fig ).

Male genitalia. Both parameres similar in length, angled ventrally, left paramere elongate and narrow, apex angled to right in ventral view, right paramere gradually narrowing to pointed apex, curved ventrally in lateral view; median lobe distinct, slightly sinuate, longer than parameres, evenly curving dorsally in lateral view, apical margin thin and fan-shaped ( Figs 12E–J View Fig ).

Comments to classification. Euglyptus iracundus ( Broun, 1893a) syn. nov. is placed as the junior synonym of E. punctatus ( Broun, 1893a) based on the examination of types by D. S. Chandler. As both species were proposed in the same paper by BROUN (1893a), the synonymy is proposed as first revisers act.

Distribution. New Zealand (North Island:AK, BP, RI, TK, WA, WN, WO, South Island: NN, SD; Fig. 16A View Fig ).