Promalactis hoenei Lvovsky, 2000

Promalactis hoenei Lvovsky, 2000

(Japanese name: Shikinami-beni-maruha-kibaga)

( Figs 5, 6 View FIGURES 1–6 , 9 View FIGURES 7–9 , 12 View FIGURES 10–12 , 15a, 15b View FIGURES 13–15 )

Promalactis hoenei Lvovsky 2000: 665 ; Wang et al. 2015: 456 View Cited Treatment .

Type locality: Shanghai ( China) .

Type depository: Museum Alexander Koenig , Bonn, Germany .

Material examined. JAPAN, Honshu: 1♂, Instisute-for-Nature-Study, Tokyo, 15.V.2017, U. Jinbo & Y. Arita leg., No. NSMT-I-L-55460 ( NSMT) ; 2♀, Akasaka-Imperial-Grounds, Tokyo, 27.V.2004, Y. Arita & M. Owada leg., No. NSMT-I-L-16988, NSMT-I-L-16992 ( NSMT) ; 2♀, Imperial-Palace, Fukiage-Gyoen , Tokyo, 14.VI.2011, Y. Arita & M. Owada leg., No. NSMT-I-L-29522, NSMT-I-L-29524 ( NSMT) ; 2♀, Imperial-Palace, Kami-Dokanbori , Tokyo, 26.V.2009, Y. Arita & M. Owada leg., No. NSMT-I-L-29509, NSMT-I-L-29510 ( NSMT) ; 2♂ 3♀, Shikinami ( N36.849111, E136.764861), Hodatsushimizu-cho , Ishikawa, 16.VI.2018, LT, S. Tomura leg., No. ST 018 GoogleMaps ♀, ST083 ♂, ST086 ♂, ST087 ♀, ST088 ♀ ( ELKU) ; 1♀, Yagoto, Nagoya-shi , Aichi, 30.V.1979, A. Nakayama leg., No. ST 198 ( NSMT) ; 1♂ 1♀, Osaka-Pref.-Univ, Sakai-shi , Osaka, 18.VI.2007, LT, T. Hirowatari, N. Hari, G. H. Huang, A. Nobuoka & S. Kobayashi leg., No. ST 173♀, ST 183♂ ( OPU) ; 1♂, same locality, 26.V.1999, N. Hirai leg., No. ST 194♂ ( OPU) ; 1♀, Mino-koen, Mino-shi , Osaka, 10.VI.2006, B.W. Lee leg., No. ST 197 ( OPU) ; 1♂, Tottori, Osaka, 25.V.2005, S. Koshino leg., No. ST 166 ( OPU) ; 2♂ 1♀, Nagakura-Pond ( N34.916944, E134.850000), Kasaishi , Hyogo, 72 m, 26.V.2015, LT, K. Sakagami leg., No. ST GoogleMaps 195♂, ST 196♂, ST 199♀ (Kota Sakagami). Kyushu : 2♂, Ikinomatsubara ( N33.583168, E130.305107), Fukuoka-shi , Fukuoka, 19.V.2020, S. Yagi leg., No. ST 228, ST229 ( ELKU) GoogleMaps ; 2♂, same locality, 28.V.2020, S. Tomura leg., No. ST 226, ST227 GoogleMaps ; 3♂ 1♀, same locality, 30.V.2020, T. Hirowatari leg., ST GoogleMaps 230♂, ST 231♂, ST 232♀, ST 233♂ ( ELKU) ; 1♂, same locality, 1.VI.2020, T. Hirowatari leg., ST 234♂ ( ELKU) .

Diagnosis. This species has the following characters: in the forewing, the three white fasciae are straight and the antemedial fascia does not reach the costal margin; in the male genitalia, the gnathos is a hemicircular band, the costa of the valva is not straight and the juxta is rectangular. These characters are shared with P. kyuparki Kim & Lee, 2016 , but the latter can be distinguished by the right sacculus with a small triangular subapical process instead of a spiniform process.

Redescription. Adult ( Figs 5, 6 View FIGURES 1–6 , 9a View FIGURES 7–9 ). Forewing length: males 5.3‒7.2 mm (n = 14) and females 5.4‒6.7 mm (n = 16). Head: vertex dark orange yellow white anteriorly; frons ocherous brown. Antennal scape white except laterally dark brown; flagellomere white basally alternated with dark brown apically. Labial palpus: second palpomere yellowish brown and yellowish white inner surface; third palpomere yellowish brown. Mesonotum and tegula dark orange yellow. Forewing dark orange yellow with three white fasciae edged with black scales; sub-basal fascia not reaching costal margin; antemedial fascia from 1/3 of posterior margin and oblique toward base; medial fascia and costal fascia linear; medial fascia from 1/2 of posterior margin and oblique toward apex; costal fascia from 2/3 of costal margin; fringe cilia orange yellow except grayish orange near tornus. Hindwing and fringe cilia gray. Hind femur and tibia pale gray; tarsus dark brown except white at the apical margin of each tarsomere.

Wing venation ( Fig. 9b View FIGURES 7–9 ). Forewing wide lanceolate, relatively broad; R 4 and R 5 stalked at 1/2; R 5 to costa; M 1 and M 2 parallel; M 2 approximate toward M 3 from anal angle of cell. Hindwing falcate: M 1 and M 2 slightly parallel; M 3 and CuA 1 stalked from anal angle of cell; 1A + 2A well developed; CuP and 3A weakly developed.

Male genitalia ( Fig. 12 View FIGURES 10–12 ). Uncus trilobed; middle lobe elongate, widened basally and apex obtuse; flanked by two lateral, small basal lobes. Gnathos belt-like, hemicircular. Valvae asymmetrical. Right valva with costa straight, gradually narrowing to a rounded apex; sacculus strongly incurved from base to subapical part; subapical part with a small, triangular process pointed inwardly; apical process spine-like, acutely upturned. Left valva with costal part narrowed basally, widening apically, extending to acutely-recurved apical process; sacculus strongly incurved, slightly narrowed subapically; apex acutely curved upwardly, larger than right apical process. Juxta rectangular, apex indented medially, about 2x length of uncus. Saccus small, triangular. Phallus broadly curved apically; vesica with single, long spiniform cornutus half length of phallus.

Female genitalia ( Figs 15a, 15b View FIGURES 13–15 ). Papilla analis narrowly-elongate, obtuse, with setae. Apophysis anterioris 2/5 length of apophysis posterioris. Ostium bursae protruded posteriorly. Lamella antevaginalis triangular, obtuse and overlaying elongated antrum. Antrum elongated, cylindrical, curved toward the right; dilated in anterior half. Ductus seminalis arising from posterior end of ductus bursae. Ductus bursae cylindrical; posterior half straight, dilated, well sclerotized and posterior end spinose; anterior half coiled, one lateral side sclerotized and narrow. Corpus bursae oval, signum absent.

Distribution. Japan (Honshu, Kyushu), China ( Fujian, Hunan, Jiangsu, Zhejiang, Shanghai).

Remarks. All individuals examined in the present study were collected in lowlands or pine forests along the coast of Honshu and Kyushu, Japan.