Leucosia anatum (Herbst, 1783)

Leucosia anatum (Herbst, 1783) View in CoL

Leucosia anatum — Hill 1982: 203, pl. 5C.

Taxonomy: Tyndale-Biscoe and George (1962), Kazmi and Tirmizi (1990), Chen and Sun (2002), Ng et al. (2014).

Type: Type material in ZMB (holotype designated by Holthuis 1959; see also Ng et al. 2014).

Type locality: Ambon, Indonesia ( Holthuis 1959; K Sakai 1999).

Distribution ( L. anatum sensu lato): Eastern Indian Ocean to West Pacific: India, Australia, Indonesia, South China, Korea and Japan ( Chen and Sun 2002).

Habitat: Subtidal; substrates of coarse sand and mud; depths 12 to 82 m ( Chen and Sun 2002).

Remarks: The form reported in Hill (1982) roughly corresponds to “morph C” in Tyndale-Biscoe and George (1962: fig. 6), and that presented by Sakai (1976: pl. 34(3)). In both of these works, past records of “ L. anatum ” in the literature had been compiled. It is apparent that this is a species complex ( Tyndale-Biscoe and George 1962; Sakai 1976; Kazmi and Tirmizi 1990; Ng et al. 2014) and needs to be revised in the future.

Seulocia latirostrata ( Shen & Chen, 1978) View in CoL #

Leucosia cf. obtusifrons View in CoL — Hill 1982: 203, pl. 5A [part?].

Seulocia latirostrata View in CoL — Wong et al. 2021: 24, fig. 32, pl. 7A.

Taxonomy: Shen and Chen (1978: as Leucosia View in CoL ),

Galil (2005b). Type: Holotype ò C-00958 and paratypes in IOASQ (now IOCAS). Type locality: Wai-luo, Guangdong, China. Distribution: East Asia: China, Taiwan and Japan ( Galil 2005b). Habitat: Subtidal; shallow estuarine seas, in soft substrates.

Seulocia rhomboidalis (De Haan, 1841) #

Leucosia rhomboidalis View in CoL — Alcock 1896: 234; Gee 1926: 160 [list];

Gordon 1931: 527; Shen 1931b: 187, text-figs. 5, 6, pl. 12(1);

Shen 1940a: 216. Leucosia maculata — Stimpson 1907: 150, pl. 18(2). Leucosides rhomboidalis — Gee 1926: 160 [list; “near Hong Kong].? Leucosia sp. — Hill 1982: 203, pl. 5B. Seulocia rhomboidalis — Galil 2005b: 51, fig. 2D; Wong et al. 2021:

24, fig. 33, pl. 7B.

Taxonomy: Chen and Sun (2002: as Leucosia ), Galil (2005b).

Type: Lectotype ò RMHN D 52261 (redesignated by Galil 2005b), and paralectotypes in RMHN (paralectotypes designated by Yamaguchi and Baba 1993; see Galil 2005b).

Type locality: Japan, locality unspecified.

Distribution: East Asia: South and East China Seas and Japan ( Chen and Sun 2002; Galil 2005b).

Habitat: Subtidal; substrate of sandy mud, soft mud or shell fragments; depths of 20 to 59 m ( Chen and Sun 2002).

Seulocia vittata (Stimpson, 1858) #

Leucosia vittata Stimpson, 1858c: 159 View in CoL ; Stimpson 1907: 149, pl. 18(3, 3a); Shen 1940a: 216 [no new specimens]; Manning and Reed 2006: 287 [“coast of China near Hong Kong ”].

? Leucosia craniolaris View in CoL — Miers 1886: 325 [part: Hong Kong material only]; Shen 1940a: 216 [not Cancer craniolaris Linnaeus, 1758 ].

Leucosides vittata — Gee 1926: 160 [list; “near Hong Kong ”].

Leucosia craniolaris View in CoL — Morton and Morton 1983: 157, fig. 9.5(5) [not C. craniolaris Linnaeus, 1758 View in CoL ].

Seulocia vittata View in CoL — Galil 2005b: 54, fig. 3B.

? Seulocia cf. vittata View in CoL — Wong et al. 2021: 24, fig. 34, pl. 7C, D.

Taxonomy: Chen and Sun (2002: as Leucosia View in CoL ), Galil (2005b).

Type: Lectotype juvenile MCZ 1351 (designated by Galil 2005b), and possible syntype in now NHMD ( Manning and Reed 2006).

Type locality: Coast of China near Hong Kong.

Distribution: Western Indian Ocean to South China Sea: from Mauritius, India, to Thailand, Malaysia, Singapore, South China, the Philippines and Indonesia ( Galil 2005b).

Habitat: Subtidal; substrates of sandy-mud, muddy sand or soft mud; depths of 11 to 41 m ( Chen and Sun 2002).

Remarks: Based on the external and G 1 morphology, L. craniolaris ( Linnaeus, 1758) was revised by Galil (2003a) and restricted to Southeast Asia, Ryukyus, eastwards in the Pacific towards Caroline and Palau Islands. Galil (2005b) reported S. vittata and S. rhomboidalis from Hong Kong. We have, however, seen substantial variation in the live colorations of “ S. vittata ” from subtidal habitats in Hong Kong (see Wong et al. 2021); more studies are needed to be done to see if the various color forms are all one species.

Urnalana haematosticta (Adams & White, 1849) #

Urnalana haematosticta View in CoL — Wong et al. 2021: 25, fig. 36, pl. 7E.

Taxonomy: Chen and Sun (2002: as Leucosia View in CoL ), Galil (2005a). Type: Syntypes NHM 1847.21 ( Galil 2005a). Type locality: “Eastern seas”. Distribution: Eastern Indian Ocean to West Pacific: from Sri Lanka, Thailand, Singapore, South China, to Indonesia, Coral Sea, New Guinea and Australia ( Galil 2005a). Habitat: Subtidal; substrates sandy-muddy or shelly; depths 24 to 100 m ( Chen and Sun 2002).