Stenohya fortuna, Xiang & Zhang & Guo, 2025

Stenohya fortuna sp. nov.

Chinese name: ỆḚÿ伪ḡ

Figs 7–12 View FIGURE 7 View FIGURE 8 View FIGURE 9 View FIGURE 10 View FIGURE 11 View FIGURE 12 .

Type material. Holotype female (Ps.- MHBU-GS2025063001 ), China: Gansu Province, Lanzhou City, Yuzhong County, Xinglong Mountain [ 35°47'53"N, 104°03'38"E], alt. 2213.08 m a.s.l., 30 June 2025, Liqi Xiang, Xiangbo Guo, Yunuo Han, Zitong Yang leg. GoogleMaps Paratypes 1female (Ps.- MHBU-GS2025063002 ), 9 protonymph (Ps.- MHBU-GS2025063003–11 ), 4 deutonymph (Ps.- MHBU-GS2025063012–15 ), same data as for holotype GoogleMaps ; 1 female (Ps.- MHBU-GS2024081101 ), China: Gansu Province, Lanzhou City, Yuzhong County, Xinglong Mountain [ 35°47'36"N, 104°05'02"E], alt. 2800.00 m a.s.l., 11 August 2024, Yancai Jin leg. GoogleMaps

Etymology. The specific epithet “fortuna ” comes from the name of the ancient Roman goddess of luck and the unpredictability of fate, in memory of the discovery of the holotype specimen while it was trapped in a psechrid spider web.

Diagnosis. Epistome triangular.Apex of pedipalpal coxa with five setae. Pedipalpal femur 5.39–6.75 as long as broad, patella 4.62–4.91 as long as broad, movable finger 1.08 times longer than hand without pedicel; fixed chelal finger with 104–111 pointed teeth.

Description. Adult female ( holotype and paratypes) ( Fig. 7A View FIGURE 7 ).

Carapace ( Figs 9E View FIGURE 9 , 10C View FIGURE 10 ). Carapace smooth, with an elevation in the middle of the eye area in lateral view, 1.12–1.22 times longer than broad, with a total of 29 setae in holotype female (34 and 36 in the two paratype females), including seven near anterior margin and six to eight near posterior margin; six to eight lyrifissures near the eyes, four lyrifissures near posterior margin; epistome small, triangular, with rounded apex; four well-developed eyes.

Chelicera ( Figs 9B View FIGURE 9 , 10B View FIGURE 10 ). Hand with seven setae and two lyrifissures, movable finger with one seta; fixed finger with 12 teeth(13 and 14 in the two paratype females), movable finger with 9 teeth; serrula exterior with 42 lamellae, serrula interior with 35 lamellae; galea developed, divided into two main branches, one branch secondarily divided into three terminal rami, while the other divided into five terminal rami ( Fig. 9D View FIGURE 9 ); rallum consisting of ten blades, all with anteriorly directed spinules, the basal-most blade shortest ( Fig. 9C View FIGURE 9 ).

Pedipalps ( Figs 9A, G View FIGURE 9 , 10A, E View FIGURE 10 ). Apex of pedipalpal coxa with five setae. Femur straight; chelal hand slender, rugby-shaped. Trochanter 1.69–2.55, femur 5.39–6.75, patella 4.62–4.91, chela with pedicel 4.19–4.78, chela without pedicel 3.94–4.46 times longer than broad, movable finger 1.08 times longer than hand without pedicel. Fixed chelal finger with eight, movable chelal finger with four trichobothria: eb and esb situated on the base of hand, grouped very closely with ib and isb; est, et and it grouped distally; ist slightly closer to est -et -ist than to isb -ib -esb - eb in fixed chelal finger; b and sb situated close to each other in basal half, st and t close to each other in distal half of movable finger. Fixed chelal finger with 104–111 pointed teeth, movable finger with 91–102 teeth, 47–68 rounded teeth at base, and 23–51 pointed ones in distal position.

Abdomen. Pleural membrane granulated. Tergites and sternites undivided, tergal chaetotaxy (I–XI): 6–7: 7–8: 8–12: 10–11: 10–14: 9–15: 11–14: 11–15: 9–15: 10–17: 7–12, sternal chaetotaxy (IV–XI): 36–40: 23–27: 25–35: 25–31: 21–31: 17–21: 13–17: 7–10, sternites VI –VIII with 7–9 medial scattered glandular setae, anal cone with two dorsal and two ventral setae. Genital area ( Figs 9F View FIGURE 9 , 10D View FIGURE 10 ): sternite II with total of 20 setae in holotype female (21 and 28 in the two paratype females), and four lyrifissures; sternite III with a row of 51 setae in holotype (50 and 54 in the two paratype females) and two lyrifissures along posterior margin .

Legs ( Figs 9I, J View FIGURE 9 , 10F, G View FIGURE 10 ). Leg I: trochanter 1.34–1.41, femur 5.14–6.68, patella 3.37–4.09, tibia 5.47–6.81, basitarsus 4.26–4.29, telotarsus 5.67–7.09 times longer than deep. Leg IV: trochanter 2.33–3.03, femur + patella 4.48–6.36, tibia 8.32–9.4, basitarsus 3.65–4.94, telotarsus 6.59–6.89 times longer than deep; tibia with two submedial tactile setae (TS =0.46–0.52, 0.88–0.97), basitarsus with two tactile setae (TS = 0.28–0.36, 0.86–0.90), telotarsus with two tactile setae (TS = 0.63–0.67, 0.24–0.28); subterminal tarsal seta bifurcate ( Fig. 9H View FIGURE 9 ). Arolium not divided, shorter than claws, claws slender and simple.

Protonymph ( paratypes).

Carapace ( Fig. 11C View FIGURE 11 ). Carapace smooth, with a slight elevation in the middle of the eye area, 1.02–1.17 times longer than broad, with a total of 19–20 setae, including four near anterior margin and six to eight near posterior margin; two lyrifissures near the eyes, two lyrifissures near posterior margin; epistome small, triangular, with rounded apex; four well-developed eyes.

Chelicera ( Fig. 11B View FIGURE 11 ). Hand with four setae and one lyrifissure, movable finger without seta; fixed finger with 9–12 teeth, movable finger with 6–7 teeth; serrula exterior with 16–20 lamellae, serrula interior with 13–15 lamellae; galea developed, divided into two branches; rallum consisting of five blades, all with anteriorly directed spinules, the basal-most blade shortest.

Pedipalps ( Figs 11A, E View FIGURE 11 ). Apex of pedipalpal coxa with two long setae. Trochanter 1.25–1.50, femur 2.63–3.36, patella 2.25–2.89, chela with pedicel 4.14–4.68, chela without pedicel 4.03–4.55 times longer than broad, movable finger 1.35–1.45 times longer than hand without pedicel. Fixed chelal finger with three, movable chelal finger with one trichobothria: eb situated on the base of hand, et distally; ist closer to eb than et; t situated on distal half of movable finger. Fixed chelal finger with 42–54 pointed teeth, movable finger with 43–50 teeth, 24–30 rounded teeth at base, and 19–20 pointed ones in distal position.

Abdomen. Pleural membrane granulated. Tergites and sternites undivided, tergal chaetotaxy (I–XI): 7–8: 6: 6: 6: 6–7: 6: 6: 5–8: 6: 5–6: 4–5, sternal chaetotaxy (IV–XI): 6: 8: 8: 8: 8–10: 8–9: 6–8: 4–6, sternites VI–VIII with 2–3 medial scattered glandular setae, anal cone with two dorsal and two ventral setae. Genital area ( Fig. 11D View FIGURE 11 ): sternite II with one seta or without seta, one lyrifissure present; sternite III with a row of six setae and one lyrifissure along posterior margin.

Legs ( Figs 11F, G View FIGURE 11 ). Leg I: trochanter 1.31–2.10, femur 3.36–4.44, patella 1.67–4.33, tibia 3.20–4.29, basitarsus 2.13–2.43, telotarsus 4.25–5.33 times longer than deep. Leg IV: trochanter 1.53–2.90, femur + patella 4.13–5.75, tibia 3.23–4.30, basitarsus 2.23–3.25, telotarsus 4.20–8.33 times longer than deep; tibia with one tactile seta (TS =0.10–0.24), basitarsus with one tactile seta (TS = 0.78–0.86), telotarsus with one tactile seta (TS = 0.45–0.60); subterminal tarsal seta bifurcate. Arolium not divided, shorter than claws, claws slender and simple.

Deutonymph ( paratypes).

Carapace ( Fig. 12 B View FIGURE 12 ). Carapace smooth, with an elevation in the middle of the eye area, 1.10–1.15 times longer than broad, with a total of 26–30 setae, including six near anterior margin and six near posterior margin; four lyrifissures near the eyes, two lyrifissures near posterior margin; epistome small, triangular, with rounded apex; four well-developed eyes.

Chelicera ( Fig. 12 C View FIGURE 12 ). Hand with five setae and one lyrifissure, movable finger with one seta; fixed finger with 6–7 teeth, movable finger with 7–8 teeth; serrula exterior with 35–39 lamellae, serrula interior with 32–34 lamellae; galea developed, divided into two main branches, one branch secondarily divided into three terminal rami, the other unbranched; rallum consisting of six blades, all with anteriorly directed spinules, the basal-most blade shortest.

Pedipalps ( Figs 12A, E View FIGURE 12 ). Apex of pedipalpal coxa with three long setae. Trochanter 1.88–2.24, femur 4.65–5.33, patella 3.52–3.79, chela with pedicel 4.73–4.91, chela without pedicel 4.55–4.77 times longer than broad, movable finger 1.35–1.41 times longer than hand without pedicel. Fixed chelal finger with six, movable chelal finger with two trichobothria: eb situated on the base of hand, very closely with ib; est, et and it grouped distally; ist situated on middle of fixed chelal finger; b situated on base of movabal finger, t distally. Fixed chelal finger with 69–72 pointed teeth, movable finger with 58–66 teeth, 32 rounded teeth at base, and 26–34 pointed ones.

Abdomen. Pleural membrane granulated. Tergites and sternites undivided, tergal chaetotaxy (I–XI): 6: 5–7: 6: 6–7: 7–8: 7–9: 10: 10: 9: 6–9: 4–6, sternal chaetotaxy (IV–XI): 6–12: 13–16: 10–16: 12–13: 12: 10–12: 10–12: 4–5, sternites VI–VIII with 3–4 medial scattered glandular setae, anal cone with two dorsal and two ventral setae. Genital area ( Fig. 12D View FIGURE 12 ): sternite II with total of 2–3 setae and two lyrifissures; sternite III with 12–13 setae and three lyrifissures along posterior margin.

Legs ( Figs 12F, G View FIGURE 12 ). Leg I: trochanter 1.64–1.77, femur 5.73–6.00, patella 3.73–4.09, tibia 3.90–4.00, basitarsus 2.60–3.25, telotarsus 3.75–4.89 times longer than deep. Leg IV: trochanter 2.00–2.47, femur + patella 5.28–7.46, tibia 6.69, basitarsus 2.82–3.00, telotarsus 4.08–5.70 times longer than deep; tibia with two tactile setae (TS =0.23– 0.37, 0.80–0.89), basitarsus with two tactile setae (TS = 0.17–0.21, 0.31–0.56), telotarsus with two tactile setae (TS = 0.20–0.31, 0.30–0.40); subterminal tarsal seta bifurcate. Arolium not divided, shorter than claws, claws slender and simple.

Measurements (in mm; length/breadth or, for legs, length/depth).

Female ( holotype and paratypes). Body length 4.52–7.30. Carapace 1.67–1.75/1.36–1.56. Pedipalpal trochanter 0.86–1.02/0.40–0.51, femur 2.31–2.48/0.36–0.46, patella 2.07–2.26/0.46–0.47, chela with pedicel 4.11–4.18/0.87– 0.98, chela without pedicel 3.86–3.92/0.87–0.98, hand without pedicel length 1.87–1.96, movable finger length 2.03–2.12. Leg I: trochanter 0.40–0.48/0.23–0.32, femur 1.06–1.27/0.19–0.22, patella 0.82–0.86/0.21–0.23, tibia 0.99–1.06/0.16–0.19, basitarsus 0.60–0.64/0.14–0.15, telotarsus 0.78–0.85/0.11–0.15. Leg IV: trochanter 0.77– 1.00/0.33–0.35, femur + patella 2.06–2.16/0.33–0.42, tibia 1.88–2.08/0.20–0.25, basitarsus 0.73–0.80/0.16–0.20, telotarsus 1.12–1.21/0.17–0.18.

Protonymph ( paratypes). Body length 1.57–2.71. Carapace 0.56–0.63/0.54–0.55. Pedipalpal trochanter 0.28– 0.35/0.20–0.28, femur 0.68–0.75/0.22–0.27, patella 0.88–0.91/0.24–0.25, chela with pedicel 2.08–2.11/0.43–0.44, chela without pedicel 0.53–0.55/0.19–0.24, hand without pedicel length 0.55–0.62, movable finger length 0.83– 0.90. Leg I: trochanter 0.17–0.21/0.10–0.13, femur 0.36–0.40/0.09–0.11, patella 0.20–0.26/0.06–0.12, tibia 0.29– 0.32/0.07–0.10, basitarsus 0.17/0.07–0.08, telotarsus 0.31–0.36/0.06–0.08. Leg IV: trochanter 0.29–0.34/0.10–0.19, femur + patella 0.65–0.69/0.12–0.16, tibia 0.41–0.43/0.10–0.13, basitarsus 0.23–0.26/0.08–0.11, telotarsus 0.40– 0.50/0.06–0.10.

Deutonymph ( paratypes). Body length 2.93–2.98. Carapace 0.87–0.85/0.74–0.79. Pedipalpal trochanter 0.45– 0.47/0.21–0.24, femur 1.07–1.12/0.21–0.23, patella 0.88–0.91/0.24–0.25, chela with pedicel 2.08–2.11/0.43–0.44, chela without pedicel 2.00–2.05/0.43–0.44, hand without pedicel length 0.83–0.87, movable finger length 1.18–1.23. Leg I: trochanter 0.23/0.13–0.14, femur 0.60–0.63/0.11–0.10, patella 0.41–0.45/0.11, tibia 0.39–0.44/0.10–0.11, basitarsus 0.26/0.08–0.10, telotarsus 0.44–0.45/0.09–0.12. Leg IV: trochanter 0.42–0.47/0.19–0.21, femur + patella 0.95–0.97/0.13–0.18, tibia 0.87/0.13, basitarsus 0.30–0.31/0.10–0.11, telotarsus 0.53–0.57/0.10–0.13.

Distribution. China: Gansu.

Remarks. Unfortunately, adult male of this new species has not been found during the field work, only adult females, deutonymphs and protonymphs were described here. The female of this new species can be distinguished from all other species, except for Stenohya harveyi Zhao, Guo & Zhang, 2025 , S. cordata Zhao, Guo & Zhang, 2025 , S. polygonia Zhao, Guo & Zhang, 2025 , S. ungulata Zhao, Guo & Zhang, 2025 , S. huangi and S. pengae , by having a rugby-shaped chelal hand. S. fortuna sp. nov. differs from these species as follows: apex of pedipalpal coxa with five long setae (seven in S. polygonia , four in S. huangi ); pedipalpal femur smooth (femur with a few tubercles prolaterally in S. ungulata ); fixed chelal finger with 104–111 pointed teeth ( 73–74 in S. harveyi , 89–93 in S. ungulata , 63–69 in S. huangi , 66–79 in S. pengae ); pedipalpal femur 5.39–6.75 times as long as broad (4.68–4.90 times in S. harveyi , 4.30–4.54 times in S. cordata , 4.57–4.63 times in S. polygonia , 4.68–5.06 times in S. ungulata ); pedipalpal patella 4.62–4.91 times as long as broad (2.93–3.23 times in S. cordata , 3.83–3.93 times in S. pengae ); pedipalpal movable finger 1.08 times longer than hand without pedicel (1.50–1.53 times in S. harveyi , 1.44–1.69 times in S. cordata , 1.49–1.59 times in S. polygonia , 1.52–1.53 times in S. ungulata , 1.26–1.38 times in S. pengae ) ( Zhao et al. 2025 Hu & Zhang 2012).