Hypselostoma khaowongense Panha, 1998

Hypselostoma khaowongense Panha, 1998

Hypselostoma khaowongensis Panha, 1998 b: 66 View in CoL , fig. 4.

Hypselostoma khaowongensis View in CoL — Hemmen and Hemmen 2001: 41.

Gyliotrachela saraburiensis Panha & Burch in Burch et al. 2003: 143–151, figs 6–7. syn. nov. View in CoL

Gyliotrachela muangon Panha & Burch in Panha et al. 2004: 67–68, fig. 7. syn. nov. View in CoL

Gyliotrachela khaowongensis View in CoL — Panha and Burch 2008: 71–72, fig. 62; Dumrongrojwattana and Wongkamhaeng 2024: 323, fig. 8.

Gyliotrachela saraburiensis View in CoL — Tongkerd et al. 2004: 143, fig. 2; Panha and Burch 2008: 78–80, fig. 68; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.

Gyliotrachela muangon View in CoL — Panha and Burch 2008: 76–77, fig. 66; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8; Gojšina et al. 2022: 137, fig. 6; Tongkerd et al. 2024: 182–184 View Cited Treatment , figs 11,13O.

Type material examined.

Thailand • holotype; S. Panha leg.; CUMZ ver. 001 • paratype of G. saraburiensis ; from the type locality; CUMZ 44047 View Materials • 2 paratypes of H. khaowongense ; from the type locality; SMF 331439 About SMF • 3 paratypes of G. saraburiensis ; from the type locality; SMF 331446 About SMF • 2 paratypes of G. muangon ; from the type locality; S. Panha leg.; SMF 331459 About SMF .

Additional material examined.

Thailand-North • 24 shells; Chiang Rai Province, Chiang Rai ca 5 km west from centre of Wat Phra That Tham Doi Kong Khao; 9°54.820'N, 99°46.642'E; 11 Feb. 2015; A. Hunyadi leg.; coll. HA GoogleMaps • 33 shells; Chiang Rai Province, ca 5 km WNW of Chiang Rai, Tham Phra cave by Mae Nam Kok river , limestone rocks near the cave entrance; 19°55.047'N, 99°47.342'E; 420 m a. s. l; Sep. 2007; A. Reischütz leg.; coll. REI GoogleMaps • 12 shells; Chiang Mai Province, 5 km northwest from Chiang Dao, vicinity of Wat Tham Chiang Dao; 19°23.628'N, 98°55.711'E; 450 m a. s. l.; 07 Feb. 2015; A. Hunyadi leg.; coll. HA GoogleMaps • 13 shells; Chiang Mai Province, Doi Pha Sao Mountain , 3 km W of Ban Prang Mao; 19°25.98'N, 99°3.48'E; 500 m a. s. l.; 19 Jun. 1987; F. G. Thompson leg.; locality code FGT-4338 ; UF 528706 GoogleMaps • 21 shells; Chiang Mai Province, NW side of Doi Pha Sam Sao ravine with Dipterocarp forest , leaf litter; 19°24.45'N, 99°2.9333'E; 800 m a. s. l.; 20 May 1988; F. G. Thompson leg.; locality code FGT-4453 ; UF 347006 GoogleMaps • 59 shells; Chiang Mai Province, NW side of Doi Pha Sam Sao ravine with Dipterocarp forest , leaf litter; 19°24.45'N, 99°2.9333'E; 800 m a. s. l.; 20. May 1988; F. G. Thompson leg.; locality code FGT-4452 ; UF 346992 GoogleMaps • 150 shells; Mae Hong Son Province, 13.9 km NW Pai , along road 1095; 19°23'N, 98°23'E; 780 m a. s. l.; 18 Mar. 1988; K. Auffenberg leg.; locality code KA-0571 ; UF 345938 GoogleMaps • 3 shells; Mae Hong Son Province, Ban Tham Lod Village , 7 km N Soppong on Road 1095; 19°34'N, 98°9'E; 790 m a. s. l.; 18 Mar. 1988; K. Auffenberg leg.; locality code KA-0574 ; UF 591328 GoogleMaps • 20 shells; Mae Hong Son Province, 21 km N Mae Hong Son, Fish Cave ; 19°26'N, 97°58'E; 380 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0593 ; UF 345672 GoogleMaps • 40 shells; Mae Hong Son Province, 40 km N of Mae Hong Son ; 19°27'N, 97°59'E; 550 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0588 ; UF 345643 GoogleMaps • 1 shell; Mae Hong Son Province, 21 km N Mae Hong Son, Fish Cave ; 19°26'N, 97°58'E; 380 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0595 A; UF 345702 GoogleMaps • 7 shells; Mae Hong Son Province, Mae Hong Son, Road 1095; 19°25'N, 97°58'E; 390 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0595 B; UF 591329 GoogleMaps • 3 shells; Mae Hong Son Province, Mae Hong Son, Road 1095; 19°25'N, 97°58'E; 390 m a. s. l.; 22 Mar. 1988; K. Auffenberg leg.; locality code KA-0595 ; UF 345699 GoogleMaps • 1 shell; Lampang Province, 6 km SW Mae Pak , limestone hill; 17°57.8833'N, 98°51.1167'E; 600 m a. s. l.; 13 June 1987; F. G. Thompson leg.; locality code FGT-4328 ; UF 591330 GoogleMaps • 11 shells; Lampang Province, limestone dome at pass 11 km SSW of Ban Pang La ; 18°27.9667'N, 99°48.3833'E; 610 m a. s. l.; 14 May 1988; F. G. Thompson leg.; locality code FGT-4441 ; UF 591331 GoogleMaps • 2 ethanol preserved shells; same locality data as previous; F. G. Thompson leg.; locality code FGT-4440 ; UF 346908 GoogleMaps • 43 shells; Phrae Province, limestone knoll 4 km N of Ban Nam Rin , deep rocks and leaves; 18°10.1333'N, 99°56.6833'E; 200 m a. s. l.; 16. May 1988; F. G. Thompson leg.; locality code FGT-4444 ; UF 346931 GoogleMaps • 1 shell; Phrae Province, Than Pha Nan Cave , 6 km NE of Rang Kwang; 18°23.1'N, 100°20.7667'E; 300 m a. s. l.; 30 Apr. 1988; F. G. Thompson leg.; locality code FGT-4400 ; UF 347403 GoogleMaps • 2 shells; Phrae Province., 4 km N of Ban Nam Rin valley with limestone ridges, leaf litter at base of boulders; 18°10.1333'N, 99°56.6833'E; 200 m a. s. l.; 16. May 1988; F. G. Thompson leg.; locality code FGT-4445 ; UF 346935 GoogleMaps • 5 ethanol preserved shells; Phrae Province, 11 km W of Phrae, on Road 1023 teak plantation and cut forest; 18°10.6833'N, 100°4.2333'E; 290 m a. s. l.; 16. May 1988; F. G. Thompson leg.; locality code FGT-4446 ; UF 346967 GoogleMaps • 19 shells; same locality data as previous; F. G. Thompson leg.; locality code FGT-4447 ; UF 591332 . Thailand-Northeast GoogleMaps • 28 shells; Petchabun Province, 24.4 km S. Petchabun on Hwy. 2275, evergreen forest in back of outcrop, leaf litter next to rocks; 16°16'N, 101°10'E; 120 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0693 ; UF 346096 GoogleMaps • 1 shell; Nakhon Ratchasima Province, 3.4 km W of Ban Mu Si ; 14.534°N, 101.380°E; 380 m a. s. l.; 05. May 1987; F. G. Thompson leg.; locality code FGT-4241 ; UF 534868 GoogleMaps • 9 shells; Nakhon Ratchasima Province, Pak Chong district, mountain above Wat Thep Phithak Punnaram ; 14°36.865'N, 101°15.926'E; 500 m a. s. l.; 04 Mar. 2023; A. Hunyadi leg.; coll. HA. Thailand-West GoogleMaps • 16 shells; Kanchanaburi Province, Erawan Nat. Park, Erawan Falls Trail; 14°22.310'N, 99°08.699'E; 85 m a. s. l.; 17 Feb. 2015; A. Hunyadi leg.; coll. HA GoogleMaps • 10 shells; Petchabun Province, 24.4 km S. Petchabun on Hwy. 2275, evergreen forest in back of outcrop, leaf litter next to rocks; 16°16'N, 101°10'E; 120 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0694 ; UF 346100 . Thailand-Central GoogleMaps • 10 shells; Phitsanulok Province, Noen Maprang district, Chomphu, Tham Phra Wang Daeng ; 16°40.651'N, 100°41.287'E; 02 Mar. 2023; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA GoogleMaps • 37 shells; Nakhon Sawan Province, 19.5 km NE intersection of Hwys 11 and 32 , limestone outcrop, base of cliff under Georissa zone; 15°14'N, 100°16'E (provided coordinates point in Chai Nat Province); 50 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0697 ; UF 346135 GoogleMaps • 1 shell; Nakhon Sawan Province, 19.5 km NE intersection of Hwys 11 and 32; 16°14'N, 100°16'E; 50 m a. s. l.; 02. May 1988; K. Auffenberg leg.; locality code KA-0697 ; UF 346126 GoogleMaps • 1 shell; Lopburi Province, Tha Wung district, Khao Samo Khon, Wat Tham Chang Pueak ; 14°54.139'N, 100°30.151'E; 30 m a. s. l.; 03 Mar. 2023; A. Hunyadi leg.; coll. HA GoogleMaps • 2 shells; Saraburi Province, behind Phra Phutthabat , limestone hill with scrubby evergreen forest, leaf litter among rocks; 14.733°N, 100.800°E; 50 m a. s. l.; 03. May 1988; K. Auffenberg leg.; locality code KA-0699 ; UF 508979 GoogleMaps . Laos • 27 shells; Luang Prabang Province, old forest on Phousi Mountain in Luang Prabang, clay, black soil, on and under rocks; 19°53.411'N, 102°08.216'E; 374 m a. s. l.; 13 Nov. 2004; A. Abdou & I. Muratov leg.; MNHN -IM-2012-25387 GoogleMaps • 6 shells; Luang Prabang Province, 3.1 km from centre of Nong Khiaw towards Pak Xeng, Tham Pha Toke , vicinity of the cave; 20°33.215'N, 102°37.722'E; 345 m a. s. l.; 05 Oct. 2019; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA GoogleMaps • 24 shells; Luang Prabang Province, 12 km from centre of Nong Khiaw towards Pak Xeng, northeast from Ban Huai Lek, left side of the road; 20°32.585'N, 102°41.161'E; 345 m a. s. l.; 05 Oct. 2019; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA GoogleMaps . Myanmar • 3 shells; Shan State, c. 3.6 km NNE of Hsihseng centre, limestone hill; 20°11′15″N, 97°16′07″E; 1040 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA GoogleMaps • 7 shells; Shan State, 9.5 km from centre of Hsihseng ; 20°14.066'N, 97°15.514'E; 1140 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA GoogleMaps • 21 shells; Shan State, 3.6 km north-northeast from centre of Hsihseng ; 20°11.248'N, 97°16.114'E; 1040 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA GoogleMaps • 63 shells; Shan State, ca 16 km from centre of Taunggyi centre towards Hopong , 1.5 km north on rd. # 4, „ Shwe Pyi Aunchonda ” monastery; 20°47.263'N, 97°8.239'E; 1110 m a. s. l.; 08 Oct. 2018; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA GoogleMaps • 57 shells; Shan State, ca 6 km east from centre of Hsihseng ; 20°8.002'N, 97°18.024'E; 500 m a. s. l.; 07 Oct. 2018; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA GoogleMaps .

Type localities.

“ A limestone hill near Takli District, Nakornsawan Province, 15°16′52″N, 100°22′38″E, 114 meters elevation ”, Thailand ( H. khaowongense ); “ East of Tepitak Tamaram Temple, Tepitak Mountain, Muang Saraburi District, Saraburi Province, 14°36'57"N, 101°15'50"E, 510 meters elevation. Thailand ” ( G. saraburiensis ); “ Muangon Cave, San Kam Pang District, Chiangmai Province, 18°47'11"N, 99°14'16"E, 280 meters elevation ” ( G. muangon ).

Differential diagnosis.

This species is similar to H. holimanae but is clearly different by the separated angular and parietal lamellae (fused in H. holimanae ) as well as the presence of numerous and variable apertural barriers (only four in H. holimanae ). Hypselostoma fruhstorferi has a narrower umbilicus and fewer apertural barriers. This species is also similar to H. cultura from which it can be separated by the less prominently shouldered last whorl and a narrower umbilicus.

Distribution.

Clearly a widespread species: its distribution stretches from Chiang Rai province in N Thailand to the southernmost known localities in Ratchaburi and Chonburi provinces. Westwards, this species stretches to Taunggyi in Myanmar and the easternmost locality is known from Luang Prabang province in Laos.

Remarks.

Examined specimens had the usual main barriers and a larger number of smaller barriers between them of very variable number (usually 2–5 between each of the main barriers). Umbilicus was always wide but sometimes more or less so (specimens from Chiang Mai ( Thailand) had the narrowest umbilicus). Last whorl is always at least very slightly keeled, in some specimens more prominent. Spiral striation is present, in some populations hardly observable and in some very rough. During comparison of the type material of G. saraburiensis and G. muangon , we have not observed any difference between these species and H. khaowongense . The former two are not compared with the latter in the respective original descriptions. Because of no difference observed, we treat both G. saraburiensis and G. muangon as junior synonyms of H. khaowongense .