Xystodesmus pallidus ( Verhoeff, 1937 ), 2025

Korsós, Zoltán & Nakamura, Yasuyuki, 2025, Review of the millipede genus Xystodesmus (Diplopoda, Polydesmida), with seven new species from the southwestern part of Japan, ZooKeys 1231, pp. 191-232 : 191-232

publication ID

https://doi.org/10.3897/zookeys.1231.141443

publication LSID

lsid:zoobank.org:pub:94B32AD5-4D03-4ECA-8B73-DB157E8AAF3E

DOI

https://doi.org/10.5281/zenodo.15013565

persistent identifier

https://treatment.plazi.org/id/60AB2267-ECD1-58F3-BE88-A3C33DCD25AE

treatment provided by

ZooKeys by Pensoft

scientific name

Xystodesmus pallidus ( Verhoeff, 1937 )
status

comb. nov.

Xystodesmus pallidus ( Verhoeff, 1937) comb. nov.

Figs 11 A – E View Figure 11 , 12 A – D View Figure 12 , 13 A, B View Figure 13

Koreoaria pallida Verhoeff, 1937: South Korea ( Verhoeff 1937: 319–320, fig. 8) View in CoL

Koreoaria pallida View in CoL : Tanabe and Shinohara 1996: 1487 (“ species possibly belonging to Xystodesmus View in CoL ”)

Koreoaria pallida View in CoL : Tanabe and Shinohara 1996: 1487

Koreoaria pallida View in CoL : Marek et al. 2014: 71

Type material examined.

Holotype: • male ( ZSMC, Reg. Nr. A 20060019 ): in 2 fragments, gonopods and legs are on separate slide ( ZSMC, Reg. Nr. A 20033594 ) (Fig. 11 View Figure 11 ) . Paratypes: • 4 females ( ZSMC, Reg. Nr. A 20060019 ) .

Description of holotype.

Male total body length ~ 21 mm; collum length 1.5 mm, width 3.3 mm; 10 th segment width 4.2 mm, metazonal length 1.1 mm. Paranotal processes from segment 6 onwards projected caudad, anterior margin broadly rounded, male 2 nd leg pair with long, slender coxal projection, 3 rd with nothing, 4 th and 5 th with small, triangular coxal processes; postgonopodal prefemoral spines strong and long, half as long as length of prefemur, sternal plate broad, coxal distance 1.2 mm, anterior-posterior sterna fused.

Male gonopods (Figs 11 A – C View Figure 11 , 12 A, B View Figure 12 ) fits well to the original description and drawing.

Female paratypes. Total body length 22–26 mm; collum length 1.6–1.8 mm, width 3.5–3.8 mm; 10 th segment paranotal width 4.4–5.1 mm, metazonal length 1.1–1.2 mm, coxal distance 1.1–1.5 mm. Gnathochilarium (Figs 12 C View Figure 12 , 13 A View Figure 13 ) and cyphopods (Fig. 13 B View Figure 13 ) are here illustrated for the first time.

Colouration (Fig. 11 D – E View Figure 11 ). Type material in alcohol completely faded. Verhoeff (1937: 319), however, provided a description which is in total agreement of Xystodesmus characters: “ braune Metatergite und rötlichgelben Längswisch auf den Seitenflügeln ” [metatergites brown, and paranota with reddish yellow longitudinal spots].

Remarks.

The flat gonopodal prefemoral process (pfp, Fig. 12 A, B View Figure 12 ) was called by Verhoeff (1937: 317, fig. 8) as “ abgespaltener Tibiotarsus ”, and in the generic comparison of Koreoaria he had set it against the prefemoral process of Pachydesmus Cook, 1895 . According to Verhoeff, the longer process made of two parts, the “ praefemorofemur ” and its disjunct ending, the solenomere ( Verhoeff 1937: fig. 8, sl). In our opinion this is a misunderstanding, as the longer process corresponds to the acropodite (Fig. 12 A, B View Figure 12 , a), and the other one is the prefemoral process (correctly interpreted by Takakuwa 1942 b for Koreoaria amoena ).

ZSMC

Zoologische Staatssammlung

Kingdom

Animalia

Phylum

Arthropoda

Class

Diplopoda

Order

Polydesmida

Family

Xystodesmidae

SubFamily

Xystodesminae

Tribe

Xystodesmini

Genus

Xystodesmus

Loc

Xystodesmus pallidus ( Verhoeff, 1937 )

Korsós, Zoltán & Nakamura, Yasuyuki 2025
2025
Loc

Koreoaria pallida

Marek PE & Tanabe T & Sierwald P 2014: 71
2014
Loc

Koreoaria pallida

Tanabe T & Shinohara K 1996: 1487
1996
Loc

Koreoaria pallida

Tanabe T & Shinohara K 1996: 1487
1996
Loc

Koreoaria pallida

Verhoeff KW 1937: 319 - 320
Koreoaria pallida Verhoeff, 1937 : South Korea ( Verhoeff 1937: 319–320 , fig. 8)
1937