Touranella Attems, 1937

Genus Touranella Attems, 1937 View in CoL

Touranella Attems, 1937: 231 (D). View in CoL

Touranella View in CoL – Attems 1938: 233 (D); Hoffman 1963: 591 (K); 1980: 172 (M); Jeekel 1968: 64 (M); Golovatch 1994: 187 (D, M); 2009 a: 6 (M, K); 2009 b: 120 (M); 2016: 139 (M, K), 2024: 408 (D); Nguyen and Sierwald 2013: 1179 (M); Golovatch and Semenyuk 2010: 125 (M); 2018: 16 (M); Nguyen et al. 2023: 171 (D, K).

Diagnosis.

Body small to medium-sized (~ 9–25 mm long, ~ 0.7–3.3 mm wide), with 20 rings. Paraterga poorly to moderately developed. Sternal lobe or cone (s) present between ♂ coxae 4. First pair of ♂ without femoral tubercles (= adenostyles). Transverse metatergal sulci distinct. Legs with neither modifications nor adenostyles. Gonopod with a long and subcylindrical coxite, slightly curved caudally; prefemoral part (= prefemorite) short, ~ 3–4 × shorter than acropodite; femoral part (= femorite) strongly reduced or very short compared to solenophore, solenomere mostly rod-shaped or subflagelliform, sheathed distally by solenophore; both lamina medialis and lamina lateralis well developed; femoral process (fp) present or absent.

Type species.

Touranella gracilis Attems, 1937 , by original designation.

Other species included.

Tournaella cattiensis Golovatch & Semenyuk, 2010 , T. champasak Nguyen, Sierwald & Ware, 2023 , T. chenla sp. nov., T. jaegeri sp. nov., T. himalayaensis Golovatch, 1994 , T. hirsuta Golovatch, 2009 , T. logunovi Golovatch, 2024 , T. moniliformis Golovatch & Semenyuk, 2018 , T. peculiaris Golovatch, 2009 , T. pilosa Golovatch, 2016 , T. srisonchaii sp. nov., and T. trichosa Golovatch & Semenyuk, 2018 .

Remarks.

This genus was originally described as monotypic, based on Touranella gracilis Attems, 1937 , which was distinguished by its markedly reduced gonopodal femoral part, the presence of a femoral process, and densely setose metaterga. The recognition of T. himalayaensis further supported these diagnostic features. However, subsequent discoveries of seven additional species in Vietnam have expanded the generic diagnosis, revealing further key variations such as the absence of a gonofemoral process and the presence of smooth metaterga ( Golovatch 2009 a, 2009 b, 2016, 2024; Golovatch and Semenyuk 2010, 2018). More recently, T. champasak from Laos was described by Nguyen et al. (2023), which included a brief discussion of the genus and its geographic distribution. The latest contribution has described another new species from southern Vietnam ( Golovatch 2024). Below a species catalogue of Touranella is presented, arranged in chronological order.