Duolandrevus ( Duolandrevus ) spinicauda Gorochov, 2016

Duolandrevus ( Duolandrevus) spinicauda Gorochov, 2016 View in CoL

( Figs 2A View FIGURE 2 , 3 View FIGURE 3 , 4 View FIGURE 4 )

Duolandrevus ( Duolandrevus) spinicauda Gorochov, 2016: 46 View in CoL .

Duolandrevus ( Duolandrevus) spinicauda gading Gorochov, 2017: 229 View in CoL

New material examined. EAST MALAYSIA; Sabah State • 1♂; Maliau Basin Conservation Area , Agathis Nature Trail; N4.70158 E116.90745, 541.9± 8.1 m.a.s.l.; 16 November 2024, 19h52; calling under bark of rotten log; coll. M.K. Tan; SBH.24.116 ( ZRC) GoogleMaps • 1♂; Maliau Basin Conservation Area , at Maliau Basin Studies Centre along Wildlife Trail; N4.73815 E116.97520, 232.9± 8.4 m.a.s.l.; 10 November 2024, 20h48; on tree trunk; coll. M.K. Tan; SBH.24.9 ( ZRC) GoogleMaps • 1♂; Maliau Basin Conservation Area , at Maliau Basin Studies Centre along Knowledge Trail (start) (24SA4); N4.74189 E116.97062, 227 m; 10–18 November 2024, night; on tree trunk; coll. T. Robillard; TR24-33 ( MNHN) GoogleMaps .

Remarks. Our male specimens are very similar to Duolandrevus ( Duolandrevus) spinicauda Gorochov, 2016 from Sarawak and Duolandrevus ( Duolandrevus) balikpapan Gorochov, 2016 from Kalimantan (both species from Borneo) in the general morphology. Our specimens show more similarities in the male FWs with D. spinicauda by the venation around the mirror area and its stridulatory file vein more sinuous; as well as more similar in the male genitalia with D. spinicauda by the pseudepiphallic lophi in profile almost not S-shaped (instead of clearly S-shaped in D. balikpapan ) (these lobes in our specimens are also shorter like in D. balikpapan ) and the pseudepiphallic paramere in profile view more rounded (distinct from both D. balikpapan and D. spinicauda ).

Our specimens are also more similar to the nominal subspecies in that the pseudepiphallic lophi curved dorsad only at the apex (instead of subapical part) and the pseudepiphallic parameres in profile more triangular (instead of flattened). Our specimens are, however, more similar to D. ( D.) spinicauda gading Gorochov, 2017 in the metanotal gland having its lateral parts of posterior keel-like fold extended more posteriorly and laterally than the nominal subspecies. These suggest that the subspecies status require revision, and more material and additional data type will be necessary.

Distribution. EAST MALAYSIA: Sarawak State: Kubah (Matang Mountains) ( type locality for nominal subspecies), Gunung Gading ( type locality for subspecies gading); Sabah State: Maliau Basin [new locality record]

Calling song ( 2♂, in the field, 25.2°C) ( Fig. 4 View FIGURE 4 ). The calling song consists of an echeme made up of an average of 24±1 syllables (21–27 syllables). The average echeme duration is 0.34± 0.03 s (0.29– 0.40 s). The first few syllables are of lower amplitudes but eventually the amplitude increases to a maximum and remains relatively consistent. The average syllable duration is of 11.1±0.6 ms (9.8–11.8 ms), the interval between consecutive syllables is 3.2±0.4 ms (2.5–3.9 ms) and the average period is 14.3±0.6 ms (13.3–15.0 ms). The frequency spectrum is pure-tonal and forms a harmonic series, with the energy peaking at a dominant and fundamental frequency of 3.66±0.19 kHz (3.47–3.84 kHz).