Delarbrea, Vieill.

DELARBREA Vieill. View in CoL Bull. Soc. Linn. Normandie 9: 342 (1865).

LECTOTYPE. — Delarbrea collina Vieill. , designated by Hutchinson, Gen. Fl. Pl. 2: 63 (1967) . Porospermum F. Muell., Fragm. 7: 94 (1870). — Type: Porospermum michieanum F. Muell.

Pseudosciadium Baill., Adansonia 12: 130 (1878), syn. nov. — Type: Pseudosciadium balansae Baill.

Monocaulous or sparsely branched, glabrous, unarmed treelets or small trees. Stem tips, petioles and base of inflorescence often glaucescent. Leaves imparipinnate, alternate, clustered at ends of branches; leaflets opposite to subopposite (or sometimes alternate, especially the lower ones), entire or remotely dentate, occasionally serrate on the first few leaves borne on new shoots (to deeply serrate or lacerate in juvenile foliage of some species), the base of the lateral leaflets oblique; rachis not articulated; petiole with an expanded, clasping base with membranous or scarious margins. Inflorescence a panicle of umbellules, terminal, erect or pendant, the lateral branches, peduncles and umbellules subtended by membranous, scarious or foliaceous bractlets, the pedicels free or basally united into groups of 2-4, articulated below the ovary. Flowers hermaphrodite and protandrous, often also functionally staminate (in andromonoecious species), actinomorphic. Sepals united below into a short tube, the 5 free lobes obtuse or triangular to rounded or oblate, valvate, sometimes with a scarious margin, not or only slightly expanding in fruit, the margins often somewhat scarious. Petals 5, imbricate or valvate, broadly ovate to obovate or spatulate, keeled within, narrowed to distinctly clawed toward the base. Stamens 5, inflexed before anthesis, the filaments stout, the anthers with 4 thecae, cream-white, dorsifixed. Ovary inferior, 2-carpellate, vestigial in staminate flowers, sometimes prolonged below into a slender stipe, surmounted by a small, rounded to depressed conic nectar disc, styles 2, free, erect and appressed at anthesis, spreading as the ± clavate stigmas become receptive. Fruit a drupe, ellipsoid-ovoid to ovoid or cylindrical, crowned by the persistent calyx and spreading styles; exocarp fleshy or spongy, the endocarp papery to bony with large oil vesicles; endosperm with shallow longitudinal grooves, not ruminate.

N OTES. — Delarbrea was last revised by LOWRY (1986a), who recognized six species, two of which were further divided into two subspecies each. Here we add a seventh species with the transfer of Pseudosciadium balansae . The genus is centered on New Caledonia, where all of the taxa are endemic except two: D. michieana (F.Muell.) F. Muell., which is restricted to Queensland, Australia, and D. paradoxa Vieill. subsp. paradoxa, which extends from New Caledonia south to Norfolk Island ( GREEN 1994) and north through Vanuatu and the Solomon Islands to the Moluccas and the Lesser Sunda Islands ( LOWRY 1986a,b, 1989; VAN BALGOOY & LOWRY 1993).

For the herbarium material cited below, geographic coordinates indicated in square brackets were assigned post facto using 1:50000 scale topographic maps of New Caledonia