Kojderus maximus Gierlasiński & Magnien, 2024

Kojderus maximus Gierlasiński & Magnien sp. nov.

( Figs 1–4 View Fig View Fig View Fig View Fig )

Material examined. HOLOTYPE: J, “ Madagaskar \ Ivohiboro Forest \ 22°34’57.5” S 46°42’25.5” E \ 30.10.2023 \ leg. A. Kepel & A. Lange ” ( MNHN) GoogleMaps . PARATYPE: ♀, “ Madagaskar \ Ivohiboro Forest, 937 m \ 22°34’59.22” S 46°42’21.18” E \ 6.11.2023 \ leg. K.Wiśniewski” ( NHC).

Diagnosis. Kojderus maximus Gierlasiński & Magnien sp. nov. is recognized by the following combination of characters: General colouration yellowish-brown; first and second antennomeres short, third antennomeres longest and slightly flattened; basal margins of hemelytra reddish- -black, inner margin of each clavus with a small, elongate black spot at the level of scutellar apex; scutellum with strongly contrasting reddish tip; lateral margins of abdomen reddish black. For additional structural characters, see the generic diagnosis.

Description. Male. Colouration, vestiture and texture: See the genus description.

Structure. Eyes: ocular index (eye width / vertex width) 0.25.Antennae: antennomere I 0.30× as long as head width, 0.50× as long as interocular distance; antennomere II 0.40× as long as head width, antennomere III longest, flattened, 1.15× as long as head width. Pronotum: maximum width of pronotum 1.75× as long as length of pronotum, lateral margins delicately serrate. Metathoracic scent efferent system with tongue-shaped peritremes, with long, narrow ostioles; mesopleural evaporatoria nearly inconspicuous, metapleural evaporatoria occupying greater part of metapleural surfaces.

Male genitalia. Pygophore ( Fig. 1D View Fig ) almost quadrangular, proctiger and parameres lying in bilobate depression of otherwise flattened upper surface, aedeagus ( Figs 3A, B View Fig ) of same construction as in other species of subfamily, fitted with two pairs of processes, dorsal one membranous, sclerotised at anteriorly pointed apex, posterolateral one short, with sclerotised apex shortly pointed; parameres ( Fig. 3C View Fig ) similar in shape to those of Haplosterna , apophysis of each slightly more broadly rounded, sensorial lobe with long setae.

Female. Mostly same as in male unless stated otherwise. Colouration. Antennomeres III and IV more reddish. Yellowish spots on hemelytra more irregular. All femora, tibiae, and tarsi yellowish-brown.

Structure. Eyes: ocular index (eye width / vertex width) 0.24. Antennae: antennomere I 0.26× as long as head width, 0.40× as long as interocular distance; antennomere II 0.36× as long as head width, antennomere III flattened, 0.82× as long as head width. Pronotum: maximum width of pronotum 1.76× as long as length of pronotum, lateral margins delicately serrate.

Female genitalia ( Fig. 4 View Fig ). Spermatheca of usual natalicoline type; apical receptacle (distal bulb) almost spherical without basal “neck”, intermediate part (pump apparatus) with two well-developed flanges; spermathecal duct bipartite, anterior duct slender, about two times shorter than posterior one, posterior duct much wider, its anterior region strongly dilated, about three times wider than narrowest part, with umbrella-shaped rod structure, rear part somewhat folded; sclerified rings visible, but only lightly contrasted.

Measurements (in millimetres). Male (n = 1) / female (n = 1). Body length: 28.00 / 34.00; width (across middle part of abdomen): 19.00 / 24.00. Head length: 2.90 / 3.20; width including compound eyes: 4.90 / 5.00; vertex width 3.00 / 3.30; eye dorsal width 0.75 / 0.80; distance between ocelli 1.30 / 1.50. Antenna. Length of antennomeres: I – 1.50 / 1.30, II – 1.90 / 1.80, III – 5.60 / 4.10, IV – 4.20 / 3.80. Pronotum length: 7.50 / 8.50; anterior width: 5.00 / 5.50; posterior pronotal width (across humeral angles) 13.00 / 15.00. Scutellum: anterior width: 8.50 / 10.00; length 6.50 / 8.00. Metafemur length: 5.60 / 6.10; metatibia length: 5.60 / 6.20; length of metatarsomeres: I – 1.40 / 1.80, II – 1.50 / 1.90.

Etymology. The name of the newly described species is the Latin adjective maximus (- a, - um), which refers to the size of this species, which is the largest of the subfamily, and also of all the Madagascan Pentatomoidea (cf. CA- CHAN 1952).

Biology. Unknown. Both specimens were collected in the rainforest ( Fig. 5 View Fig ).

Distribution. Kojderus maximus Gierlasiński & Magnien sp. nov. is known from two close localities in Ivohiboro Forest, in south-central Madagascar.