Chaerilobuthus complexus Lourenço and Beigel, 2011

Chaerilobuthus complexus Lourenço and Beigel, 2011

( Figs 11–16 View Figure 11 View Figure 12 View Figure 13 View Figure 14 View Figure 15 View Figure 16 , 39 View Figure 39 )

Chaerilobuthus complexus Lourenço and Beigel, 2011: 635–639 , figs 1–11.

Type material: Holotype: 1 juv. (sex unknown) (Senckenberg Museum, Frankfurt, Germany), Cretaceous (Cenomanian) burmite: MYANMAR: Kachin State (precise locality unknown) .

Diagnosis: The following combination of characters is diagnostic for Chaerilobuthus complexus : carapace anteromedian margin aligned with anterolateral margins ( Figs 11A View Figure 11 , 14A View Figure 14 ); carapace anterior margin slightly depressed submedially ( Figs 12A View Figure 12 , 15A View Figure 15 ); lateral ocelli protruding ( Figs 12A View Figure 12 , 15A View Figure 15 ); sternum relatively narrow ( Figs 17B View Figure 17 , 20B View Figure 20 ); pedipalp chela robust, ratio of chela length: chela manus width less than 5 ( Figs 13C View Figure 13 , 16 View Figure 16 ); pedipalp chela fixed finger median denticle row comprising c. seven oblique subrows ( Figs 13C View Figure 13 , 15E View Figure 15 ); pectines each with five teeth ( Figs 12B View Figure 12 , 15D View Figure 15 ); mesosoma relatively wide, tergite VII width much greater than length ( Fig 11E View Figure 11 ); metasomal segment I width much greater than length ( Fig. 12I, J View Figure 12 ).

Description: Based on the material examined ( NIGP 200655, 201160).

Carapace: Anterior carapace margin with few, short microsetae; anteromedian depression narrow, shallow; anterosubmedial depressions absent; lateral ocular carinae absent; posteromedian and posteromarginal sulci obsolete; surfaces almost smooth to sparsely covered in microsetae ( Figs 12A View Figure 12 , 15A View Figure 15 ). Median ocular tubercle situated anteromedially; median ocelli absent; one pair of large posterolateral major lateral ocelli present.

Coxosternal region: Surface smooth, sparsely covered in macrosetae ( Figs 12B View Figure 12 , 39D View Figure 39 ). Leg I maxillary lobes (coxapophyses) distal margins slightly curved, dilate anteriorly and adorned with serrula comprising c. 14–16 spinules ( Fig. 39D View Figure 39 ). Sternum pentagonal; lateral margins sublinear, parallel; ventral surface flat, without distinct concave region or median sulcus; posteromedian depression wide, shallow; posterior margin slightly recurved ( Figs 12B View Figure 12 , 15B View Figure 15 ).

Chelicerae: Fixed finger and manus, prolateral and ventral surfaces each with numerous long, dense macrosetae ( Fig. 12C, G View Figure 12 ). Moveable finger markedly overlapping fixed finger ( Fig. 12D View Figure 12 ) such that cheliceral dentition partly visible; moveable finger with long dorsal distal (dd) denticle, small dorsal subdistal denticle, stout dorsal median denticle and long ventral distal (vd) denticle; fixed finger with long distal denticle and small subdistal denticle ( Figs 18E, F View Figure 18 , 20C View Figure 20 ). Moveable finger with prominent serrula, comprising moderately long spinules in proximal three-quarters ( Fig. 12C, G View Figure 12 ).

Pedipalps: Segments moderately robust; surfaces sparsely covered by short microsetae and few macrosetae ( Figs 11 View Figure 11 , 13 View Figure 13 , 14 View Figure 14 , 16 View Figure 16 ). Femur with three carinae evident; prodorsal and proventral carinae distinct, serrate, prodorsal carinae with small apophysis proximally ( Figs 13A View Figure 13 , 15F View Figure 15 ). Patella with four carinae evident; prolateral surface, dorsoventral ‘vaulted’ projection (‘anterior process’) distinct with prominent pair of apophyses or tubercles (‘patellar spurs’), each possessing one macroseta; retrodorsal and retroventral carinae smooth, costate ( Figs 13B View Figure 13 , 15I View Figure 15 ). Chela relatively robust; manus globose; chela with six carinae evident; digital carina incomplete, distinct on fixed finger, obsolete on manus; dorsomedian carina reduced to few granules proximally on manus; prodorsal carina distinct, comprising several strong spiniform granules; promedian and proventral carinae obsolete, each comprising few isolated granules; retroventral carina obsolete, smooth, and costate ( Figs 13C, D View Figure 13 , 15H View Figure 15 ); dorsal secondary, subdigital, retromedian and ventromedian carinae absent or obsolete. Fixed and moveable fingers median denticle rows each comprising seven oblique, slightly imbricate subrows, separated by large serrate prolateral and retrolateral denticles ( Figs 13C View Figure 13 , 15E View Figure 15 ). Trichobothrial pattern Type D, β configuration. Pedipalp femur with 10 trichobothria visible, five d, two i, and three e trichobothria, trichobothria d 1, d 2, d 4, and d 6 noticeably smaller than others (‘petite’); trichobothrium e 1 situated proximal to trichobothrium d 3; trichobothrium e 3 vertically aligned with trichobothrium d 6 ( Figs 13A View Figure 13 , 16 View Figure 16 ). Patella with four trichobothria visible, including three d and one i trichobothria; trichobothrium i 1 situated distal to trichobothrium d 3 ( Figs 13B View Figure 13 , 16 View Figure 16 ). Chela with eight trichobothria visible; fixed finger with two d, two e and one i trichobothria; manus with two E and one V trichobothria; trichobothrium Et 1 situated distally on manus, almost aligned with or proximal to moveable finger condyle; trichobothrium eb situated on fixed finger, slightly distal to moveable finger condyle; trichobothrium esb situated medially on fixed finger; trichobothrium V 2 situated distally on manus and near retroventral carina ( Figs 13C, D View Figure 13 , 16 View Figure 16 ).

Legs: Tibia lateral and ventral surfaces each with scattered macrosetae ( Figs 12K, L View Figure 12 , 15G View Figure 15 ). Basitarsi each with pair of spinule rows, comprising seven or eight relatively long spinules, distally ( Figs 12H, L View Figure 12 , 15G View Figure 15 ). Ungues moderately long, distinctly curved; dactyl pronounced, pointed ( Figs 12K, L View Figure 12 , 15G View Figure 15 ).

Pectines: Basal piece long, narrow ( Figs 12B View Figure 12 , 15D View Figure 15 ). Three marginal lamellae and four median lamellae; surface sparsely covered in macrosetae; longitudinal suture between marginal and median lamellae distinct. Fulcra absent or obsolete. Teeth rounded, distal half of each tooth occupied by sensilla field; peg sensilla very long, columnar with distinct pore.

Genital operculum: Two opercular sclerites narrow and completely separated, as long as pectinal plate ( Fig. 12B View Figure 12 ).

Mesosoma : Pretergites surfaces almost smooth ( Fig. 12E View Figure 12 ); post-tergites I– VI surfaces finely granular, acarinate ( Fig. 12E View Figure 12 ); tergite VII surface sparsely granular with four moderately developed carinae (paired dorsosubmedian and dorsolateral carinae) in posterior half of segment ( Fig. 11A View Figure 11 ). Sternites surfaces smooth, with scattered macrosetae; posterior margins broadly recurved; respiratory spiracles (stigmata) small, ovoid, oblique ( Figs 12F View Figure 12 , 15C View Figure 15 ).

Metasoma: Segments I– III progressively increasing in length, I wider than long, others longer than wide ( Fig. 12I, J View Figure 12 ). Segments I– III lateral and ventral surfaces each with few, short macrosetae; dorsal surfaces each with distinct depression. Segments I– III each with six distinct carinae (paired dorsosubmedian, dorsolateral and ventrolateral carinae); dorsosubmedian carinae distinct, markedly serrate; dorsolateral carinae distinct, moderately serrate; ventrolateral carinae obsolete on segment I, moderately developed on II and III.

Material examined: MYANMAR: Kachin State: Noije Bum near Hukawng Valley, precise locality unknown, Cretaceous (Cenomanian) burmite: one juv. (sex unknown), mesosomal segments IV, V and telson absent ( NIGP 200655 View Materials ), one juv. (sex unknown), sinistral pedipalp absent ( NIGP 201160 View Materials ) .