Tomosvaryella seticosta Földvari, Skevington & Motamedinia, 2023

Tomosvaryella seticosta Földvari, Skevington & Motamedinia sp. nov.

urn:lsid:zoobank.org:act:45103086-5720-4F92-A13F-0471C6A4967E

Figs 89A–E View FIGURE 89 , 139 View FIGURE 139 , 155B View FIGURE 155

Diagnosis: This species can be recognized by the upper side of the basal costal cell having two long, brown bristles and 3–4 short, light brown bristles; symmetrical surstyli, broadened basally, moderately narrowed to apex in dorsal view ( Fig. 89A View FIGURE 89 ); gonopods with two different different-sized finger-like projections towards surstyli ( Fig. 89B View FIGURE 89 ); hypandrial apodeme distinct; subepandrial sclerite with a long protrusion along its middle in ventral view ( Fig. 89B View FIGURE 89 ).

Description: MALE: Body length: 2.8–3.2 mm. Head. Flagellum short acuminate; yellow-brown. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 1.5 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1/3, but it is silver all along the edge of the eyes. Thorax. Postpronotum pale, with 8–10 pale bristles. Mesonotum (viewed obliquely from front) brownish pollinose, anterior part more grey, grayish also from the side and the posterior edge (2 small patches). Scutellum silvery pollinose and with 6–8 weak bristles along the distal edge. Dorsocentral bristles distinct, dark, somewhat longer along frontal edge, numerous dark bristles around postpronotum present. Halter, knob pale with brownish tinge, stem brown. Legs. Mid coxa with 4–5 long brown bristles. Trochanters and femora dark brown, shining ventrally, silvery pollinose posteriorly except shiny hind femur; knees and basal 1/4 of tibiae yellow (tibiae otherwise brown), tarsal segments yellow-brown, ventrally paler, last segment darker. Hind trochanter with a half circle, no distinct bristles, half circle with flat part dorsally; two ventrobasal spines present on fore femur but absent on mid femur. Ventroapical row of spines on fore femur 5–7; 8–10 small, black spines on mid femur; hind femur without ventroapical spines, except 18–20 equally spaced bristles posteroventrally, longer on distal half, longest up to 1/2 the width of hind tibia at distal end, outer edge (lateral side) with 10–12 stronger dark spines in the middle line. Subapical (distal) spines on first four tibiae absent. Hind metatarsus moderately flattened and almost as long as 2–5 combined, with scrub-like bristles ventrally, strong spine-like bristles dorsally on all tarsi. Pulvilli shorter than last tarsal segment. Wing. Length: 3.5–3.6 mm. Upper side of basal costal cell with two long brown bristles and 3–4 short light brown bristles. Fourth costal section 2–2.5 times as long as third costal section. Cross-vein r-m at middle of discal cell. 4–5 distinct brown setulae on tegula. Abdomen. Viewed obliquely from front tergites subshining black, tergite 1 silvery grey, silvery pollinose lateral spots on tergite 5. Dispersed strong dark bristles on all tergites absent. Lateral spines on first tergite present, 5–7 short dark bristles up to 0.6 times as long as width of hind femur at base. Postabdomen in dorsal view: T6, S7 invisible; T5 3 times as long as ST8. Genitalia without dissection: ST8 small, rounded in dorsal view, brown and without distinct bristles, but with velvet-like coverage; membranous area round, mostly directed posteroventrally; epandrium brown, surstyli yellow and brown. Genitalia. Genital capsule in dorsal view: epandrium slightly longer than wide (MLE:MWE = 1.07). Surstyli rather symmetrical, broadened in basal two thirds, then narrowed to apex ( Fig. 89A View FIGURE 89 ). Genital capsule in ventral view: gonopods small with two finger-like projections towards surstyli, right is longer than left one, subepandrial sclerite with a protrusion along its middle; hypandrial apodeme prolapsed phallic guide strong ( Fig. 89B View FIGURE 89 ); Genital capsule in lateral view: both surstyli widened, curved towards sternite ( Fig. 89D–E View FIGURE 89 ). Ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 89C View FIGURE 89 ).

FEMALE: Body length: 3.1–3.2 mm. As male except for the following characters. Frons, eyes separated, as wide as 1.2–1.4 times the width at antennae; completely silver-grey pollinose, only shiny black directly in front of ocellar triangle, ca. 1/6 of the frons, very short transition zone; frons with a short ridge in front of the ocelli in the median line. Enlarged ommatidia silvery shining. Pulvilli and claws about 2–2.5 times as long as last tarsal segment, smaller on hind tarsomeres. Female abdomen with silver spots on sides of tergite 6. Ovipositor. Straight in ventral view, robust, curved towards sternite apically in lateral view (yellow piercer, brown base), reaching distal end of 3 rd segment; base brownish pollinose with occasional bristles. LP:LB = 3.5. LDP:LPP = 2.5. ( Fig. 155B View FIGURE 155 ).

Etymology: The name refers to the bristles found on the basal costal cell and comes from the Latin seta, bristle.

Examined material: HOLOTYPE: AUSTRALIA: Queensland: Ridgepole Waterhole, 24 km ESE of Musselbrook Resource Centre, Lawn Hill Nat [ional] P[ar]k, 18°40’S, 138°22’E, 180m, 13.V.1995, G. Daniels, M.A. Schneider, JSS8283 (1♂, QM) GoogleMaps ; PARATYPES: AUSTRALIA: Queensland: Bulburin State Forest , 24°32’S, 151°34’E, 27–29. V GoogleMaps .1960, F.A. Perkins, JSS8257 (1♂, QM) ; 200 km west Windorah , 25°25’S, 142°39’E, Malaise trap in dry creek bed, 4.IX.1997, S. Winterton, J.&A. Skevington, JSS8325 (1♀, QM) GoogleMaps ; Ridgepole Waterhole, 24 km ESE of Musselbrook Resource Centre, Lawn Hill National Park , 18°40’S, 138°22’E, 180m, 5. V GoogleMaps .1995, G. Daniels & M.A. Schneider, JSS8299 (1♀, QM) ; Carnarvon Station near Homestead , 24°48’S, 147°45’E, 750m, Eucalyptus woodland , 12.X.2014, Lambkin, Wright, J. & K. Wilson, hand net, CNC576417 View Materials (1♂, CNC) GoogleMaps ; Western Australia: Karijini National Park, Juna Downs Road , 22°40’S, 118°26’E, 797m, in open mulga with flowering Ptilotus , 25.IV–14. V GoogleMaps .2003, C. Lambkin & T . Weir , Malaise trap, JSS15991 (1♂, ANIC) ; New South Wales: Culgoa National Park, 8km West-Northwest Cawwell Homestead, Diemunga Lagoon ( CGN2 View Materials M), 29°3’S, 146°60’E, Coolibah, 30.I– 18. V GoogleMaps .2010, C. Lambkin, R . Olsen & B. Shieban, Malaise trap, CNC575120 View Materials (1♂, CNC) ; Northern Territory: Darwin , 12°27’S, 130°50’E, 6–9. XII GoogleMaps .1963; 8–9. XII GoogleMaps .1963, J. Sedlacek , CNCD5571 , CNCD5575 (2♂, BPBM) .

Distribution: Australia (New South Wales, Northern Territory, Queensland, Western Australia) ( Fig. 139 View FIGURE 139 ).

Notes: This species is widespread but rarely collected.