Tomosvaryella pterygia Földvari, Skevington & Motamedinia, 2023

Tomosvaryella pterygia Földvari, Skevington & Motamedinia sp. nov.

urn:lsid:zoobank.org:act:6ABB3BF2-0EBE-48D6-AB4C-6D9F60FD7342

Figs 83A–E View FIGURE 83 , 136 View FIGURE 136 , 152F View FIGURE 152 , 156E View FIGURE 156

Diagnosis: This species can be recognized by the hind trochanter bearing 6–7 bristles ventrally; surstyli rectangular-shaped, pointing towards each other with distinct bristles on lateral distal corners directed laterally in dorsal view ( Fig. 83A View FIGURE 83 ), phallic guide with 3–4 dorsolateral spines, subepandrial sclerite long with fine transverse streaks ( Fig. 83B View FIGURE 83 ).

Description: MALE: Body length: 2.2–2.4 mm. Head. Flagellum acuminate, yellow-brown. Face silvery pollinose. Frons, upper part shining black, lower part silvery pollinose; eyes touching for distance equal to 1.5–2 times ocellar triangle. Occiput silvery pollinose, slightly less so on upper half. Thorax. Postpronotum pale brown, with 2–3 whitish bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part silvery pollinose, also silvery from the side. Scutellum grey pollinose, without bristles. Dorsocentral bristles weakly developed, pale, somewhat longer on frontal part. Halter, knob pale, stem dark black. Legs. Trochanters and base of femora brown, femora dark brown, shining ventrally (hind femur also posteriorly), knees, basal 1/6 of tibiae, tarsal segments yellow, last segment dorsally brown. Mid coxa with three strong bristles at inner apical corner. Ventroapical row of spines on fore femur missing, 4–6 very short, black spines on mid femur; no spines on hind femur, only 3–4 flattened thick bristles on keel. Subapical (distal) spines on first four tibiae absent. Hind trochanter with 6–7 short dark bristles on ventral side, basal spines absent (short pale basal bristles on fore and mid femur may be present). Hind tarsomeres slightly flattened, hind metatarsus slightly longer than 2–4 combined. Pulvilli slightly shorter than last tarsal segment. Wing. Length: 2.4–2.5 mm. Upper side of basal costal cell with one long distinct bristle ( Fig. 152F View FIGURE 152 ). Fourth costal section 2.5–3 times as long as third costal section. Cross-vein r-m at middle of discal cell. Setulae on tegula absent. Abdomen. Viewed obliquely from front tergites brownish pollinose, hind margin silvery grey on tergite 1, sides completely silvery on tergites 4 and 5. Dispersed short dark bristles absent. Lateral bristles on first tergite: 2–3, pale brown, as long as width of hind femur at base. Postabdomen in dorsal view: T6, S7 invisible; T5 as long as ST8. Genitalia without dissection: ST8 appears large, mainly dark brown and forming a right-angle corner (90 degrees) in lateral view at the lower end of membranous area; membranous area slit-like, vertical, distal part of epandrium paler and surstylus tips whitish, surstyli relatively small. Genitalia. Genital capsule in dorsal view: surstyli rectangular-shaped with acute tips on apical inner corners, epandrium wider than long ( Fig. 83A View FIGURE 83 ), 0.5 times as long as ST8; cerci symmetrically placed, small; ST8 with slit-like membranous area. Genital capsule in ventral view: Subepandrial sclerite covered with horizontal transverse streaks; gonopods symmetrical; phallic guide with a membranous projection to the side; phallus trifid, branches short; phallic guide with 3–4 dorsolateral spines, ejaculatory apodeme flat; sperm pump elongated, linear ( Fig. 83B View FIGURE 83 ). Genital capsule in lateral view: both surstyli straight, bent before apex ( Fig. 83D–E View FIGURE 83 ).

FEMALE: Body length: 2.2–2.6 mm. As male except for the following characters. Frons, eyes separated, as wide as 2 times the width at antennae; completely silver-grey pollinose until 1/6–1/4 from the antennae and along median line up to 2/3 of the frons, less so along the edges, smooth transition from grey pollinose to shiny black. Enlarged ommatidia silvery shining. Pulvilli and claws about 1–1.5 times as long as last tarsal segment on metatarsi. Ovipositor. Base of ovipositor dark brown with some scattered short dark bristles, piercer slightly down curved. LP: LB = 1.7. LDP:LPP = 3.0. ( Fig. 156E View FIGURE 156 ).

Etymology: Named after the Greek word pterygion for fin, in reference to the fin-shaped keel on the hind femur of males.

Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Yathong Nature Reserve near 8 mile tank, 32°31’S, 145°37’E, 218m, across wide wash: field of flowers: open woodland, 4–25.x.2003, C. Lambkin & N. Starick, Malaise trap, CNCD3560 (1♂, QM); PARATYPES: AUSTRALIA: New South Wales: Culgoa National Park, 13.5km SW Cawwell Homestead (CGN1M), 29°7’S, 146°56’E, Callitris , 30.I–18.V.2010, QM Team, B. Shiebaan & R. Ohlsen, Malaise trap, 19325, CNC576619, CNC576624, CNC576626, CNC576629 (4♂, QM); Culgoa National Park, 4.5km WNW Cawwell Homestead, Claypan (CGN3M), 29°3’S, 147°1’E, 192m, Gidgee, 30.I–18.V.2010, B. Schiebaan, Malaise trap, 17220, CNC576533–35 (3♂, QM); Culgoa National Park, 8km WNW Cawwell Homestead, Diemunga Lagoon (CGN2M), 29°3’S, 146°60’E, Coolibah, 22.XI–21.XII.2009, C. Lambkin, B. Shiebaan & N. Starick, Malaise trap, DB 9 (2nd bottle), 19302, CNC593683–84 (2♂, QM); Culgoa National Park, 8km West–Northwest Cawwell Homestead, Diemunga Lagoon (CGN2M), 29°3’S, 146°60’E, Coolibah, 30.I– 18.V.2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, 19322, CNC575125 (1♂, QM); Ledknapper National Reserve, 5km N Beulah Homestead (LDN1M), 28°19’S, 147°10’E, Allocasuarina , 18.III–15.V.2010, S. O’Sullivan & C. Lambkin, Malaise trap, 12 DB, 19361, CNC576716 (1♂, QM); Warrumbungles National Park, Woolshed at Wombelong Creek, 31°16’S, 148°57’E, 420m, rocky valley, cedar forest, 12–16.XII.1995, M.E. Irwin, Malaise trap, JSS14028 (1♂, INHS); Culgoa National Park, 8km WNW Cawwell Homestead, Diemunga Lagoon (CGN2M), 29°3’S, 146°60’E, Coolibah, 22.XI–21.XII.2009, C. Lambkin, B. Shiebaan & N. Starick, Malaise trap, CNC593677 (1♂, CNC); Culgoa National Park, 13.5km SW Cawwell Homestead (CGN1M), 29°7’S, 146°56’E, Callitrus, 21.XII.2009 – 30.I.2010, B. Schiebaan, Malaise trap, CNC576570, CNC576578, CNC576580, CNC576588 (4♂, QM); Blackbox, 22.XI–21.XII.2009, C. Lambkin, B. Shiebaan, N. Starick, Malaise trap, 11 DB, 19300, CNC576655, CNC576658, CNC576664 (3♂, QM); Callitris , 30.I–18.V.2010, QM Team, B. Shiebaan & R. Ohlsen, Malaise trap, 19325, CNC576606, CNC576610, CNC576639, CNC576641 (4♂, QM); Culgoa National Park, 8km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN2M), 29°3’S, 146°60’E, Coolibah, 30.I–18.V.2010, C. Lambkin, R. Olsen, B. Shieban, Malaise trap, 19322, CNC575128, CNC575132, CNC575136 (3♂, QM); Billabong Creek, Conargo, 35°19’S, 145°9’E, 23–30.iv.1978, Z. Liepa, JSS9170 (1♂, ANIC); Culgoa National Park, 1km North-Northwest Cawwell Homestead, Claypan (CGN4M), 29°4’S, 147°3’E, 136m, Gidgee, 30.i.2010, B. Schiebaan, Malaise trap, CNC575350 (1♂, QM); Culgoa National Park, 8km West-Northwest Cawwell Homestead, Diemunga Lagoon (CGN2M), 29°3’S, 146°60’E, Coolibah, 30.i–18.v.2010, C. Lambkin, R. Olsen & B. Shieban, Malaise trap, CNC575142, CNC575144, CNC575147, CNC575151–4 (7♂, QM); Culgoa National Park, 8km West–Northwest of Cawwell Homestead, Diemunga Lagoon (CGN2M), 29°3’S, 146°1’E, 136m, Coolibah, 21.xii.2009 – 30.i.2010, B. Schiebaan, Malaise trap, CNC574755–6 (2♂, QM); Fowlers Gap Research Station, 31°5’S, 141°42’E, 29.xi–2.xii.1981, I.D. Naumann & J.C. Cardale, ex. ethanol via cellosolve-xylene, JSS9234–6 (3♂, ANIC); Jerilderie, 35°22’S, 145°43’E, 28.xi.1978, J. McGechan, JSS8739 (1♂, ASCU); Mulwaree Ponds, Inveralochy Bridge, near Yass, 34°44’S 149°45’E, 27.ii.1966, Z. Liepa, JSS9238 (1♂, ANIC); Warrumbungle National Park, 31°14’S, 149°1’E, 19.x–3.xi.1997, J. Skevington & S. Winterton, Malaise trap, JSS8436 (1♀, QM); Yathong Nature Reserve near Triangle Tank, 32°38’S, 145°33’E, 200m, in sandy wash in low closed mallee, 3– 25.x.2003, C. Lambkin & N. Starick, Malaise trap, JSS16038 (1♂, ANIC); Yathong Nature Reserve near 8 Mile Tank, 32°38’S, 145°41’E, ~ 205m, field of flowers: open woodland, 3–25.x.2003, C. Lambkin & N. Starick, Malaise trap; CNCD3618, CNCD3634 (2♂, QM); Yathong Nature Reserve near 8 mile tank, 32°31’S, 145°37’E, ~ 218m, across wide wash: field of flowers: open woodland, 4–25.x.2003, C. Lambkin & N. Starick, Malaise trap, CNCD3559, CNCD3563 (2♂, QM); Queensland: 2.5km West–Northwest of Plevna Downs Homestead (PD3 P2), 26°40’S, 142°35’E, 133m, Gidgee, 25.XI–17.XII.2008, R. Mackenzie, pitfall trap, 17309, CNC575273 (1♂, QM); Culgoa Floodplain National Park, 11.8km N Headquarters (CG2AM), 28°54’S, 147°8’E, 151m, Brigalow, 20.I–18.III.2010, C. Kelly & A. Coward, Malaise trap, 19271, CNC576676, CNC576678 (2♂, QM); Elizabeth Creek, Boodjamulla NP, 18°14’S, 138°5’E, 170m, Bloodwood open, 18–22.IV.2005, M. Mathieson & G. Smith, Malaise trap, 12398, JSS50564 (1♂, CNC); Kurumba, 17°29’S, 140°53’E, 18.IV.1983, J.F. Donaldson, JSS8957 (1♂, QDPC); Plevna Downs, 2.3km SSW Arima (PD2 M), 26°32’S, 142°31’E, 134m, Gidgee, 25.XI–17.XII.2008, R. Mackenzie, Malaise trap, 17292, CNC595264 (1♂, QM); Plevna Downs, 2.5km WNW Homestead (PD3), 26°40’S, 142°35’E, 133m, Gidgee, 16.IX–2.X.2008, Starick, Lambkin & Mackenzie, Malaise trap, 16272, CNC591861 (1♂, QM); 12.7km South-Southeast of Plevna Downs Homestead (PD8 M), 26°47’S, 142°39’E, 145m, sandy creek bank, 25.XI–18.XII.2008, R. Mackenzie, Malaise trap, 17341, CNC593802 (1♂, QM); Culgoa Floodplain National Park Headquarters, 11.8km N, 28°55’S, 147°8’E, 150m, Brigalow, 9.XII.2007 – 16.I.2008, N. Starick, C. Lambkin & R. Raven, Malaise trap, 15683, CNC591849 (1♂, QM); Culgoa Floodplain National Park, 11.8km N Headquarters (CG2AM), 28°54’S, 147°8’E, 151m, Brigalow, 20.I–18.III.2010, C. Kelly, A. Coward, Malaise trap, 19271, CNC576687–90, CNC576700 (5♂, QM); 18.XI–17.XII.2009, C. Lambkin, A. Coward, C. Kelly & N. Starick, Malaise trap, CNC576510, CNC576512 (2♂, QM); Culgoa Floodplain National Park, 11.8km North of Headquarters (CG2AM), 28°54’S, 147°8’E, 151m, Brigalow Black box, 17.XII.2009 – 20.I.2010, C. Kelly & A. Coward, Malaise trap, CNC574522 (1♂, QM); Currawinya National Park, 10 mile bore, 0.5km Northwest, 28°52’S, 144°28’E, 130m, Eucalyptus , 13.X–15.XII.2007, C. Lambkin & N. Starick, Malaise trap, CNC575118 (1♂, QM); Currawinya National Park, 15.1km Southwest of Headquarters (CW9M), 28°55’S, 144°26’E, 131m, Eremophila , 12–26.IX.2008, Lambkin, Starick, Townsend, Malaise trap, 17241, CNC574663 (1♂, QM); Plevna Downs, Tompilly Hill base (PD6 M), 26°44’S, 142°39’E, 187m, Gidgee, 24.IV–24.VII.2008, G. Turner & R. Mackenzie, Malaise trap, CNC595284 (1♂, QM); 2.5km West-Northwest of Plevna Downs Homestead (PD3 P2), 26°40’S, 142°35’E, 133m, Gidgee, 25.xi–17.xii.2008, R. Mackenzie, pitfall trap, CNC575271–2 (2♂, QM); 200 km west Windorah, 25°25’S, 142°39’E, ~Malaise in dry creek bed, 4.ix.1997, S. Winterton, J&A. Skevington, JSS8326–9 (3♂, 1♀, QM); 204 km west Windorah, 25°42’S, 140°53’E, Malaise in dry creek bed, 4.ix.1997, S. Winterton, J. & A. Skevington, JSS8338 (1♂, QM); 91.2 km west Windorah, 25°22’S, 141°47’E, Malaise in dry creek bed, 31.viii–5.ix.1997, S. Winterton, J. & A. Skevington, JSS8342 (1♂, QM); Charleville, 26°24’S, 146°14’E, 15.ii.1970, A.L. Dyce, car net, ex. spirit via ethyl acetate, JSS8619, JSS8623 (2♂,ANIC); Kroombit Tops National Park, hilltop, 24°27’N, 150°56’E, 22.xii.2014, J.H., A.M. & A.W. Skevington, CNC384505–8 (4♂, CNC); Mount Moffatt summit, 25°4’S, 148°3’E, 1097m, 18.xi.2014, J.H., A.M. & A.W. Skevington, CNC374868, CNC374963, CNC375031, CNC375036 (4♂, CNC); Plevna Downs, Tompilly Hill base (PD6M), 26°44’S, 142°39’E, 187m, Gidgee, 24.vii–17.ix.2008, N. Starick, C. Lambkin & R. Mackenzie, Malaise trap, CNC574713 (1♂, QM); Ridgepole Waterhole, 24 km ESE of Musselbrook Resource Centre, Lawn Hill National Park, 18°40’S, 138°22’E, 7.v.1995, G. Daniels & M.A. Schneider, JSS8300 (1♀, QM); South Australia: 6 km South Maynard’s Bore, Everard Pk. Station, 34°57’S, 138°34’E, by beating Grevillea , 6.XI.1970, G. Gross, JSS8708 (1♂, SAM); Leigh Creek, 30°29’S, 138°25’E, 15.IX.1972, Z. Liepa, JSS9206 (1♂, ANIC); Near Victory Well, Everard Park Station, 27°3’S, 132°30’E, 2–4.XI.1970, G. Gross, Malaise trap, JSS8710 (1♂, SAM); Gammon Ranges National Park, south end of Weetootla Gorge, 30°29’S, 139°16’E, 537m, hilltop, 11.X.1997, J. Skevington & C. Lambkin, [hand collected, thorax and 2 legs remaining, rest used for sequencing], JSS1259 (1♂, QM); JSM10829, JSS1261 (1♂, QM); Flinder’s Ranges National Park, Brachina Gorge, Heysen Hilltop , 31°20’S, 138°33’E, 9.x.1997, J.&A. Skevington & C. Lambkin, JSS8379–81, JSS8389–90, JSS8393 (6♂, QM); Gammon Ranges National Park, Weetootla Gorge, 30°28’S, 139°13’E, 537m, 13.x.1997, J. Skevington & C. Lambkin, JSS8370, JSS8372–4 (4♂, QM); Gammon Ranges National Park, south end of Weetootla Gorge, 30°29’S, 139°16’E, 537m, hilltop, 11.x.1997, J. Skevington & C. Lambkin, hand collected, JSS4425 (1♂, QM), hilltop, 11.x.1997, J. Skevington & C. Lambkin, JSS8345, JSS8347–8, JSS8352–3, JSS8356–7, JSS8361, JSS8363–4, JSS8366–7 (12♂, QM); Koonamore, inside KVR, 32°3’S 139°23’E, salt bush sweep, 18.ii.1974, P. Greenslade, [genitalia missing; head damaged], JSS8818 (1♂, SAM); Near Victory Well, Everard Park Station, 27°3’S, 132°30’E, 2–4.xi.1970, G. Gross, Malaise trap, JSS8713 (1♂, SAM); 2.xi.1970, G.F. Gross, Malaise trap, JSS8711–2 (2♂, SAM); Swan Reach Conservation Park, 34°35’S, 139°29’E, [in mallee], 29.ix–4.x.1997, S. Winterton, J.&A. Skevington & C. Lambkin, Malaise trap, JSS8420 (1♂, QM); Western Australia: 70km S of Newman; Great Northern Highway, 23°54’S, 119°45’E, 626m, near trees in small wash, 6–18.V.2003, M.E. Irwin & F.D. Parker, Malaise trap, CNCD6902 (1♂, CNC); 48 km NW Carnarvon; Blowholes Road; sandy trail, 24°35’S, 113°31’E, dry vegetated coastal dunes, 27.iv–11.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15827 (1♂, CNC); 11 km E Marble Bar at Brockman Creek, 21°9’S, 119°52’E, ~ 187m, damp, sandy creek bed, 2–14.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15683, JSS15685 (2♂, CNC); 137 km S Newman, 30 km W Kumarina Road House, 24°28’S, 119°40’E, 21–23.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, 19.iv.–23.v.2003, JSS16417–9 (1♂, 2♀, CNC); 15 km N Auski Road House, 22°16’S, 118°46’E, 402m, water hole in Acacia woodlands, 3–16.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15846 (1♀, CNC); 158 km S Newman, 9 km N Kumarina Road House, 24°38’S, 117°37’E,~ 638m, in wide sandy wash, 18–21.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS16406, JSS16408, JSS16410–4 (4♂, 2♀, CNC; 1♀, ANIC); wide sandy wash, 24.iv–7.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15652, JSS15705, JSS15707–8, JSS15710–4 (8♂, 1♀, CNC); ~ 638m, wide sandy wash, 7–18.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15719, JSS15721, JSS15724–5, JSS15727–31 (8♂, 1♀, CNC); 171 km east of Marble Bar, 21°17’S, 121°14’E, 300m, dry wash with flowers, 2– 15.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15873 (1♀, CNC); 30 km ESE Three Rivers Station, dry bed of Gascoyne River, 25°14’S, 118°57’E, 504m, dry river bed, 24.iv–7.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15651 (1♀, CNC); 47 km S Pardoo Road House on Shay Gap Road, 20°23’S 120°1’E, 170m, dry wash near Spinifex , 1–14.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15701 (1♂, CNC); 70 km S Newman on Great Northern Highway, 23°54’S, 119°45’E, 626m, small wash with Eucalyptus , 24.iv–6.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15882 (1♀, CNC); Karijini National Park, Banjima Drive, 10 km W visitors center, 22°29’S, 118°22’E, 743m, 4–16.v.2003, M.E. Irwin & F.D. Parker, JSS15823 (1♂, CNC); Karijini National Park, Hamersley-Mount Bruce Road, 22°34’S, 118°18’E, 757m, dry rocky creek bed, Eucalyptus grassland, 25.iv– 14.v.2003, C. Lambkin & T. Weir, Malaise trap, JSS16119–20, JSS16122 (3♂, ANIC); Karijini National Park, Juna Downs Road, 22°40’S, 118°26’E; 22°42’S, 118°25’E, ~ 797m, 789m, in open mulga with flowering Ptilotus ; between drying pools in rocky Turee Creek, 19–25.iv.2003, C. Lambkin & T. Weir, Malaise trap, JSS16171, JSS16255, JSS16286, JSS16288, JSS16290–1, JSS16293–4, JSS16297, JSS16299 (6♂, 4♀, ANIC); 22°40’S, 118°26’E, 22°44’S, 118°25’E, 797m, 798m, in open mulga with flowering Ptilotus , dry Turee Creek, grassy open Eucalyptus scrub, 25.iv–14.v.2003, C. Lambkin & T.Weir, Malaise trap, JSS15993, JSS15997, JSS16001, JSS16004, JSS16088, JSS16095, JSS16307, JSS16309–11 (6♂, 4♀, ANIC); Karijini National Park, Karijini Drive , 22°34’S, 118°18’E, 814m, open Eucalyptus grassland, 25.iv–14.v.2003, C. Lambkin & T. Weir, Malaise trap, JSS15585 (1♀, ANIC); Karijini National Park, Weano Gorge Road, 22°22’S, 118°15’E, 22°24’S, 118°15’E, 775m, 718m, on hilltop, in open Eucalyptus , 25.iv–14.v.2003, C. Lambkin & T. Weir, Malaise trap, JSS16145, JSS16218, JSS16232– 4, JSS16237–9 (5♂, 3♀, ANIC); Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22°42’S, 118°25’E, 789m, 25.iv–14.v.2003, C. Lambkin & T. Weir, Malaise trap, CNCD3822 (1♂, CNC); Kimberley, Lennard River Crossing, Gibb River Road, 17°23’S, 124°44’E, ~in riverine thicket, 14–28.vii.1988, T.F. Houston, Malaise trap, JSS8734–5 (1♂, 1♀, WAM); Lane Poole Reserve, 32°49’S, 116°6’E, 315m, Banksia grandis / Jarrah woodland on bauxite ridge, 7–13.xi.2003, C. Lambkin & J. Recsei, Malaise trap, JSS16064 (1♂, ANIC); Mount Augustus National Park, south slope on Saddle Trail, 24°18’S, 116°48’E, 400m, small dry wooded wash, 25.iv–7.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15886 (1♀, CNC); Millstream-Chichester National Park, Black Hill Pool, 21°20’S, 117°15’E, 176m, over wide dry rocky creek bed, Eucalyptus , 7–12.v.2003, C. Lambkin & D. Yeates, Malaise trap, JSS16313 (1♂, ANIC); Millstream, 21°35’S, 117°4’E, 10.iv.1971, D.H. Colless, JSS9198– 9 (2♂, ANIC); 12.iv.1971, D.H. Colless, JSS9192–3, JSS9200 (3♂, ANIC); ~ 3 km north of Denham, 25°55’S, 113°32’E, hilltop, 12.xii.1999, J. Skevington, JSS7319, JSS7343, JSS7360, JSS7365, JSS7380, JSS7387, JSS7396, JSS7407 (8♂, QM); 23km E. Marble Bar, 21°11’S, 119°57’E, 222m, Sandy wash, 15.v.2003, F.D. Parker, M.E. Irwin, pan trap, CSIRO–SCHLINGER PILBARA EXPEDITION 19.iv.–23.v.2003, CNCD6951, CNCD6953–4, CNCD6956, CNCD6958–9 (2♂, 4♀, ANIC); Lochada, 1.2km East-Southeast of Bolada Camp, 29°12’S, 116°32’E, 282m, open York Gum woodland, Hakea, 15–18.IX.2009, Lambkin, Monteith, Malaise trap, CNC595382–3 (2♂, CNC); Australia: Rhodes grass, 19.I.1966, accession#D.P.; R2367; DC.HC.NY.D.L., JSS8971 (1♂, QDPC).

Distribution: Australia (New South Wales, Queensland, South Australia, Western Australia) ( Fig. 136 View FIGURE 136 ).

Notes: This is one of the most common species of Tomosvaryella in Australia and is widespread, mostly in the outback. Based on the shape of the genitalia, this species is close to T. latipoda sp. nov.. It differs by having rectangular-shaped surstyli, its apex is wider and covered with dense long bristles in dorsal view ( Fig. 83A View FIGURE 83 ), but the surstyli of T. latipoda sp. nov. is narrowed at the apex with long bristles before the apex ( Fig. 53A View FIGURE 53 ). Intraspecific genetic distance ranges from 0.0% to 2.5%. This species is genetically most similar to T. kooncheraensis sp. nov. (1.1 % pairwise divergence) (Supplementary file 3).