Tomosvaryella leimonias ( Perkins, 1905 )

Tomosvaryella leimonias ( Perkins, 1905) View in CoL

Figs 56A–E View FIGURE 56 , 129 View FIGURE 129 , 150F View FIGURE 150

Pipunculus leimonias Perkins, 1905 View in CoL

Diagnosis: This species can be recognized by the mid tibia with one distinct and long apical spur, as long as the mid metatarsus ( Fig. 150F View FIGURE 150 ); surstyli elongated, apical margin flattened in dorsal view ( Fig. 56A View FIGURE 56 ); both gonopods extended towards surstyli with a small triangular projection in its middle; phallic guide with 3–6 dorso-dorsolateral spines before apex in ventral view ( Fig. 56B View FIGURE 56 ).

Redescription: MALE: Body length: 2.6 mm. Head. Scape, pedicel and arista drak brown. Pedicel with 2–3 upper bristles; flagellum brown, gray pollinose and tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance as long as the length of ocellar triangle. Occiput grey pollinose covered by scattered short dark bristles. Thorax. Postpronotum light yellow, with 4–5 long brown bristles in upper margin. Mesonotum (viewed obliquely from front) gray pollinose; scutum with scattered dark bristles at anterior supra-alar area, extended to posterior and uniseriate rows of intra-alar same sized dark bristles. Postalar callus dark brown with 2–3 long, as long as bristles in scutum, light brown bristles in the middle. Scutellum silvery pollinose and with 25–30 long, as long as scutum bristles, dark bristles, scattered throughout it. Halter, knob pale, stem light brown, base dark. Legs. Mid coxa with 3–4 long dark brown bristles. Coxae, trochanters, femora dark, all femora shining ventrally, knees and basal 1/3 of tibiae yellow (tibiae otherwise dark brown), tarsomeres brown ventrally, dark dorsally. Mid trochanter with 4–8 short dark bristles ventrally. Hind trochanter silvery pollinose covered by some (10–15) scattered short brown bristles ventrally; two ventrobasal spines absent on fore mid femur. Hind tibia with two rows of short brown bristle on anterior and posterior side. Mid tibia with one distinct and long apical spur, as long as mid metatarsus ( Fig. 150F View FIGURE 150 ). Hind tarsomeres slightly flattened to one side, covered by short dark brown bristles, hind metatarsus as long as 2–4 combined; pulvilli shorter than last tarsal segment. Wing. Length: 2.7 mm. Upper side of basal costal cell with one long and two short dark brown bristles. Fourth costal section 1.5–2 times as long as third costal section. Cross-vein r-m at middle of discal cell. 2–3 short dark setulae on tegula. Abdomen. Tergites 1 silvery pollinose with 15–17 different-sized lateral bristles; tergite 2–5 brown pollinose, covered by scattered short dark bristles. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE:MWE = 1.6). Surstyli symmetrical, left surstylus longer than right one, broadened at base, then strongly constricted, elongated to apex, apical margin flattened ( Fig. 56A View FIGURE 56 ). Genital capsule in ventral view: both gonopods equal in height, extended towards surstyli with a small triangular projection in its middle; phallic guide with 3–6 dorso-dorsolateral spines before apex, one is longer than others; subepandrial sclerite small ( Fig. 56B View FIGURE 56 ); Genital capsule in lateral view: both surstyli curved to sternite, widened in the middle ( Fig. 56D–E View FIGURE 56 ); ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 56C View FIGURE 56 ).

FEMALE: unknown.

Examined material: HOLOTYPE: AUSTRALIA: Queensland: Brisbane , 27°28’S, 153°1’E, VI.1904, GoogleMaps

JSS4150 (1♂, ZMAN) ; Non-type material examined: New South Wales: Nelligen , 35°38’S, 150°8’E, (tidal flat) GoogleMaps ,

1.II.1973, D.H. Colless, JSS8499 (1♂, ANIC). Distribution: Australia (New South Wales, Queensland) ( Fig. 129 View FIGURE 129 ). Notes: It appears that Perkins only based this species on one specimen, so this is the holotype. The New South

Wales specimen was taken on a tidal flat so efforts to find this species should likely focus on mangrove habitats.