Villaria purpurea (Elmer) Arriola & Alejandro, 2020

3. Villaria purpurea (Elmer) Arriola & Alejandro View in CoL , comb. nov.

Tricalysia purpurea Elmer (1906) View in CoL 70. — Hypobathrum purpureum (Elmer) Merr. (1923) View in CoL 534. — Type: Elmer 7659 (lost in PNH, no duplicates found), Philippines, Province of Tayabas, Mount Banahaw. Neotype (designated here): FB (Borden) 2928 (neo K!; isoneo K!, L), Philippines, Province of Bataan, Mount Mariveles. See note 2.

Shrubs to small trees, up to 3 m tall; branches glabrous. Stipules triangular, 5–7 by 3–4 mm, keel prominent from the base to the tip, outer surface glabrous, inner surface pilose at the base, margin sparsely ciliate, colleters numerous, 0.8–1 mm long. Leaves: petioles 7–8 by 1.5–2 mm, glabrous; blades lanceolate to narrowly ovate, 7–12 by 2.5–4 cm, base acute, apex acuminate, subcoriaceous, glabrous on both sides; secondary veins 8–12 on each side. Inflorescences supra-axillary, 3.5–5 mm above the node, few- to many-flowered dichasial cymes; peduncle 3–4.5 by c. 0.9 mm, glabrous; pedicels 1–1.3 cm long, glabrous; bracts triangular, 0.3–0.8 by c. 0.3 mm, apex acuminate and sharply pointed, glabrous outside, densely pilose inside, margins sparsely ciliate; colleters numerous at the base inside. Calyx View in CoL : tube infundibuliform, 2.5–5 by c. 3 mm, glabrous throughout; lobes ovate, 0.9–1.3 by 0.5–1 mm, glabrous on both sides, margin sparsely ciliate. Corolla View in CoL : tube infundibuliform, 1.5–2 mm long, glabrous on both sides; lobes ovate, 3–3.5 by 2–2.5 mm, glabrous outside, sparsely pilose on the base inside, margin sparsely ciliate. Anthers 2.5–3 by c. 1 mm, base truncate, apex obtuse. Style 1–2.3 mm long; stigmatic branches elliptic, 4–4.5 by c. 1 mm, papillate inside. Fruits green, purple when mature, 2.4–4 by c. 2 cm; stalk 3.5–4.6 cm long, glabrous. Seeds angular-ovoid, 6 – 8 mm long, greenish to yellow.

Distribution — Philippines ( Bataan, Quezon, Aurora, Sorsogon, Surigao, Visayas, Zambales Prov.).

Habitat & Ecology — In primary forest to deep wooded ravines, also near coastal areas. Flowering: April to June; fruiting: May to September.

Additional specimens studied. THE PHILIPPINES, Alabat, Quezon, BS (Ramos & Edaño) 48148 (P); Alas-asin, Bataan, Arriola & Banag TA002 (USTH); Angat, Bulacan, BS (Ramos & Edaño) 34067 (P); Baler, Aurora, Alejandro et al. 10­110 (USTH); Bataan, FB (Pascual) 28779 (P); Bulusan, Elmer 15925 (L); Ilocos, BS (Ramos) 33007 (K); Irosin, Elmer 16598 (K); Lamao, Bataan, Borden 2928 (P), Borden 21726 (K); Maria Aurora, Arriola & Alejandro 11­048 (USTH); Montalban, Rizal, Loher 6428 (K); Quezon, Merrill 34067 (P); Rizal, BS (Ramos) 41871 (P); Surigao, Banag & Tandang SU024 (PNH); Visayas, Langenberger 2074 (L).

Notes — 1. The lanceolate to narrowly ovate leaf shape of V. purpurea resembles that of V. acutifolia and V. glomerata . However, V. purpurea differs from all other known Villaria species by having the longest fruit stalks of all.

2. Elmer (1906) indicated Elmer 7959 to be the type. No duplicate with this number was found. In the Biodiversity Heritage Library, the number in the scan of Elmer (1906) has been changed by hand to 7616 (a number, among others, also mentioned by Merrill 1923), of which a duplicate is present in E. However, it is not clear that Elmer made a typing error. Therefore, we select a neotype here.