Foordana distincta, Haddad, 2025

Haddad, Charles R., 2025, And they just keep coming: four new genera of dark sac spiders from southern Africa (Araneae, Trachelidae), African Invertebrates 66 (1), pp. 19-64 : 19-64

publication ID

https://doi.org/10.3897/AfrInvertebr.66.139299

publication LSID

lsid:zoobank.org:pub:2D20D0B7-7DF4-42E6-A70C-EB1875BCCD26

DOI

https://doi.org/10.5281/zenodo.14726721

persistent identifier

https://treatment.plazi.org/id/7F85D5E9-9210-52EC-9E5B-8FE38A5C2374

treatment provided by

African Invertebrates by Pensoft

scientific name

Foordana distincta
status

sp. nov.

Foordana distincta sp. nov.

Figs 1 A – C View Figure 1 , 2 View Figure 2 , 3 View Figure 3 , 4 View Figure 4 , 5 View Figure 5 , 6 View Figure 6 , 7 View Figure 7 , 8 View Figure 8 , 9 D, G, H View Figure 9

Material examined.

Holotype. South Africa • ♂; Free State Province; Clocolan, Amahelo-ho-Spitskop ; 28°48.561'S, 27°39.255'E; 17 Mar. 2010; C. Haddad leg.; base of grass tussocks; NCA 2010/349 . GoogleMaps

Paratypes. South Africa • 4 ♀; together with holotype GoogleMaps Eastern Cape Province; 1 ♀; Queenstown / Komani district, Farm Arphen ; 31°51.546'S, 26°34.112'E; 1160 m a. s. l.; 4 Apr. 2021; C. Haddad leg.; hand collecting; NCA 2021/1044 GoogleMaps 1 ♂ 2 ♀; Amatola Mountains, Hogsback, Amatole Forestry Company ; 32°33.727'S, 26°54.924'E; 1460 m a. s. l.; 23 Mar. 2013; C. Haddad leg.; active search, grass tussocks and fynbos in grassland; NCA 2014/321 GoogleMaps Free State Province: 1 ♂ 2 ♀; Clocolan, Amahelo-ho-Spitskop ; 28°48'S, 27°39'E; 9 Mar. 2007; C. Haddad leg.; Rhus litter; NCA 2007/1334 GoogleMaps 2 ♀; same collection data as for preceding; 8 Mar. 2007; dense grass; NCA 2007/1336 GoogleMaps 1 ♀; same collection data as for preceding; grassy litter near dam; NCA 2008/2897 GoogleMaps 1 ♀; same collection data as for preceding; Eucalyptus litter; NCA 2007/1335 GoogleMaps 1 ♀; Clocolan, Amahelo-ho-Spitskop ; 28°48.561'S, 27°39.255'E; 2010; A. Jones leg.; in garden and around house; NCA 2010/328 GoogleMaps 3 ♀; Wepener district, Farm Dereham , 29°52.671'S, 27°04.456'E; 1520 m a. s. l.; 31 Mar. 2024; C. Haddad leg.; hand collecting in grassland; NMBA 19614 View Materials GoogleMaps KwaZulu-Natal Province; 3 ♀; Pietermaritzburg, Town Bush ; 29°33'S, 30°21'E; 15 Apr. 1976; F. Wanless & A. Russell-Smith leg.; ground layer in grassland; BMNH GoogleMaps Western Cape Province: 1 ♀; Houw Hoek, Houw Hoek Inn ; -34.2046, 19.1537; 275 m a. s. l.; 25 Nov. 2021; C. Haddad leg.; hand collecting in fynbos; NMBA 18796 View Materials GoogleMaps .

Diagnosis.

The male of this species shares with F. flavipoda sp. nov. a simple curved embolus in ventral view (Figs 8 B View Figure 8 , 10 A View Figure 10 ), whereas it is S-shaped with a distal loop in F. kasouga sp. nov. (Fig. 11 B View Figure 11 ). Furthermore, it can be recognised by the generally similar size of the dorsal and ventral retrolateral tibial apophyses in lateral view (Fig. 8 C View Figure 8 ), whereas the ventral is far larger than the dorsal in F. flavipoda sp. nov. (Fig. 10 B View Figure 10 ) and the dorsal is larger than the ventral in F. kasouga sp. nov. (Fig. 11 C View Figure 11 ). Females can be separated from F. flavipoda sp. nov. and an undescribed species from Zimbabwe by the absence of a median hood (compare Fig. 9 G – J View Figure 9 ).

Description.

Male (holotype, Amahelo-ho-Spitskop, NCA 2010/349 ): Measurements: CL 2.36, CW 2.03, AL 2.77, AW 1.87, TL 5.15, FL 0.14, SL 1.33, SW 1.15, AME - AME 0.07, AME - ALE 0.04, ALE - ALE 0.41, PME - PME 0.11, PME - PLE 0.19, PLE - PLE 0.68, MOQ: AW 0.35, PW 0.36, L 0.40. Length of leg segments: I 2.05 + 0.95 + 1.61 + 1.22 + 0.81 = 6.64; II 1.80 + 0.88 + 1.35 + 1.12 + 0.76 = 5.91; III 1.32 + 0.63 + 0.88 + 1.07 + 0.43 = 4.33; IV 1.92 + 0.85 + 1.60 + 1.78 + 0.63 = 6.78. Carapace deep red-brown, with four pairs of mediolateral mottled markings, corresponding to palps and first three pairs of legs (Fig. 2 A View Figure 2 ); surface finely wrinkled, appearing asetose but with very short sparse setae (Fig. 3 B View Figure 3 ); fovea short, distinct, at ⅔ CL. AER slightly procurved, almost straight; clypeus height equal to AME diameter; AME very slightly larger than ALE; AME separated by distance equal ½ their diameter; AME separated from ALE by distance equal to slightly less than ⅓ AME diameter; PER slightly recurved, PLE slightly larger than PME; PME separated by distance equal to slightly more than ⅘ their diameter; PME separated from PLE by distance equal to 1 ½ PME diameter. Chelicerae, endites and labium dark orange-brown; anterior surface of cheliceral paturon covered with fairly dense long, fine setae mesally, sparse laterally; promargin with three slightly separated teeth, median tooth largest; retromargin with three adjacent teeth on common base, decreasing in size distally; endites slightly paler. Sternum dark orange, with faint black mottled patches laterally, border dark orange-brown; surface smooth, with scattered erect setae. Abdomen elongate-oval, creamy-grey dorsally, with narrow yellow-brown dorsal scutum, leaving narrow sliver between scutum and black lateral markings (Fig. 2 A View Figure 2 ); dorsum with continuous black line from anterior to posterior, with pair of narrow lines in posterior half of abdomen, joined by six faint transverse chevron markings; sides black (Fig. 2 B View Figure 2 ); venter creamy-grey, with pair of black mediolateral lines from epigastric fold to spinnerets (Fig. 2 C View Figure 2 ). Legs I to IV dark yellow-orange, I darker than others; ventral cusps present on tibiae, metatarsi and tarsi I and II (Fig. 8 A View Figure 8 ). Palp (Figs 7 A, B View Figure 7 , 8 B, C View Figure 8 , 9 D View Figure 9 ) dark red-brown; tegulum oval, with embolus originating mesally, initially with broad base directed prolaterally, bending sharply before gradually curving to tip, directed at 2 o’clock; vRTA slender, subtriangular, with rounded tip pointed inwards in ventral view; dRTA slightly smaller with sharp tip, triangular in lateral view.

Female (paratype, Amahelo-ho-Spitskop, NCA 2010/349 ): Measurements: CL 2.48, CW 2.05, AL 3.58, AW 2.35, TL 5.60, FL 0.24, SL 1.32, SW 1.05, AME - AME 0.06, AME - ALE 0.03, ALE - ALE 0.37, PME - PME 0.13, PME - PLE 0.14, PLE - PLE 0.65, MOQ: AW 0.31, PW 0.40, L 0.35. Length of leg segments: I 2.12 + 1.02 + 1.52 + 1.20 + 0.86 = 6.72; II 1.90 + 0.92 + 1.35 + 1.02 + 0.80 = 5.99; III 1.47 + 0.77 + 1.00 + 1.12 + 0.55 = 4.91; IV 2.25 + 0.98 + 1.80 + 1.95 + 0.70 = 7.68. Carapace deep orange-brown, with three paired mediolateral mottled grey markings and mottled patch in front of fovea (Fig. 2 F View Figure 2 ); AER slightly procurved, almost straight; clypeus height equal to ALE diameter; AME slightly larger than ALE; AME separated by distance approximately ⅖ their diameter; AME separated from ALE by distance equal to ⅕ AME diameter; PER slightly recurved, PME and PLE subequal; PME separated by distance equal to their diameter; PME separated from PLE by distance equal to PME diameter. Chelicerae, endites and labium orange-brown; cheliceral promargin with three slightly separated teeth, median tooth largest; retromargin with three adjacent teeth on common base, decreasing in size distally. Sternum deep yellow-brown, borders darker, with mottled lateral markings near coxal bases; surface finely wrinkled, covered with scattered short, fine setae. Abdomen without scutum, dorsal, lateral and ventral markings as for male (Fig. 2 F – H View Figure 2 ). Legs I to IV creamy-yellow, tibiae, metatarsi and tarsi I and II yellow-brown. Epigyne (Figs 7 F View Figure 7 , 8 D, E View Figure 8 , 9 G, H View Figure 9 ) with broad, sharply curved atria mesally, with copulatory openings entering short lateral copulatory ducts before entering large teardrop-shaped anterior ST II, each with oval lateral accessory gland; connecting ducts exiting ST II posteromesally, curving laterally, making a twist before entering bilobed posterolateral ST I, with short channel leading to fertilization ducts.

Etymology.

The species name is the Latin for “ distinct, clear ”, referring to the well-defined abdominal markings of this species.

Distribution.

Widespread across the southern half of South Africa (Fig. 12 View Figure 12 ).

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Trachelidae

Genus

Foordana