Ursinia glandulosa Magee & Boatwr., 2014

2. Ursinia glandulosa Magee & Boatwr. View in CoL , sp. nov.

Type:— SOUTH AFRICA. Northern Cape, Vioolsdrif (2817): Richtersveld, Rosyntiesberg (–AC), 30 August 1977, Oliver, Tölken & Venter 297 (holotype PRE!) .

Woody perennial shrublet, 0.30–0.40 m tall; densely branched, branches stout, with stalked glands. Leaves cauline, alternate, 8.0–12.0 × 4.0–7.0 mm, trifid to bi-trifid, glandular-punctate, white felty when young, becoming glabrescent; stalked gland present; lobes narrowly ovate to lanceolate, 1.0–5.0 × ± 1.0 mm, mucronate; mucro acute. Capitula radiate, heterogamous, ca. 15–20 mm in diam., solitary; peduncle 50–80 mm long at anthesis, sparsely pilose. Involucre hemispherical, 5.0–7.0 × 8.0–10.0 mm, white felty; involucre bracts 5- to 6-seriate, brownish along inner margins; outer bracts narrowly ovate, 1.5–2.0 mm long, margins scarious; middle bracts narrowly oblong, 4.0– 4.5 mm long, margins scarious; inner bracts narrowly oblong, 5.0–7.0 mm long, apical appendages broadly ovate, obtuse. Receptacle paleate. Paleae scarious, truncate, repand to slightly erosulate. Ray florets neuter, ca. 12 to 16; lamina 4–6 mm long, yellow. Disc florets bisexual, numerous, yellow, 5-lobed; lobes cucullate, with sessile glandular trichomes. Anthers slightly sagittate at base; apical appendages cordate. Style terete; branches linear, truncate, sweeping hairs apicallydorsally. Pappus uniseriate, scales 5. Cypselas unknown.

Distribution and ecology: — Ursinia glandulosa is an endemic species of the Richtersveld ( Fig. 3 View FIGURE 3 ) where it is found in two major vegetation types, i.e. SKr 8 Rosyntjiesberg Succulent Shrubland (here at high elevations, 1060 m on Rosyntjiesberg Quartzite) and SKr1 Central Richtersveld Mountain Shrurbland ( Mucina et al. 2006).

Diagnostic characters: — Ursinia glandulosa , as the name implies, differs from all other currently recognised Ursinia species by the conspicuous stalked glands on the leaves and branches ( Fig. 1P View FIGURE 1 ). This species has previously been confused with U. punctata Brown (1887: 670) with which it shares the woody perennial habit, weakly pinnatisect leaves, apical appendages only on the outer involucral bracts and the truncate paleae. It differs further from the latter in the short stout branches ( Fig. 1M View FIGURE 1 ), the densely white felty young leaves and involucral bracts ( Fig. 1N, O View FIGURE 1 ) and the presence of apical appendages ( Fig. 2E View FIGURE 2 ) only on the innermost involucral bracts (branches lanky, young leaves sparsely pilose, involucral bracts glabrescent to sparsely pilose and both inner two-most involucral bracts apically appendaged in U. punctata ).

Additional specimens examined: — SOUTH AFRICA. Northern Cape Province: 2817 (Vioolsdrif): Armmanshoek (–AC), September 1995, Williamson & Williamson 5777 ( NBG) .