Heterogomphus, Burmeister, 1847

Key to the adult males of Heterogomphus View in CoL species from Colombia

Modified from Ratcliffe et al. (2020)

The absence of developed cephalic or pronotal structures is not a constant character that allows the differentiation of females and males because minor males may appear similar to females. However, males have the last abdominal ventrite medially emarginate ( Fig. 10A View FIGURE 10 ), while females have the last abdominal ventrite completely arcuate, not emarginate ( Fig. 10B View FIGURE 10 ).

1. Elytra completely, densely rugopunctate or with punctate rows................................................. 2

1´. Elytra smooth........................................................................................ 7

2. Elytra with long, moderately dense to dense setae............................................................ 3

2´. Elytra glabrous or only with short, sparse setae on apex....................................................... 4

3. Clypeal anterior margin truncate ( Fig. 7B View FIGURE 7 ); pronotal punctures small ( Fig. 7F View FIGURE 7 ).................................................................................... H. kogui Neita Moreno, Clavijo-Bustos, & Ratcliffe , new species

3´. Clypeal anterior margin broad, deeply and triangularly emarginate ( Fig. 7A View FIGURE 7 ); pronotal punctures large ( Fig. 7E View FIGURE 7 )........................................................................................... H. carayoni Dechambre View in CoL

4. Clypeal apex truncate................................................................... H. curvicornis Prell View in CoL

4´. Clypeal apex broad, deeply and triangularly emarginate....................................................... 5

5. Color dark reddish brown; elytra finely punctate............................................... H. rugicollis Prell View in CoL

5´. Color black; elytra densely punctate or rugopunctate......................................................... 6

6. Mesotibial apex with 2–3 large teeth; parameres as in Fig. 3L–M......................... H View in CoL View FIGURE 3 . schoenherri Burmeister

6´. Mesotibial apex with 1 large tooth; parameres as in Fig. 3H–I.............................. H View in CoL View FIGURE 3 . mniszechi (Thomson)

7. Clypeal apex truncate.................................................................................. 8

7´. Clypeal apex broadly emarginate......................................................................... 9

8. Mesotibia and metatibia with truncate apex ( Fig. 10C View FIGURE 10 ).................................... H. dilaticollis Burmeister View in CoL

8´. Mesotibia and metatibia at apex with two large teeth ( Fig. 10D View FIGURE 10 )............................ H. telamon (Burmeister) View in CoL

9. Apex of the medial trough in the elevated section of the pronotum broad, wider than width of head, broadly and deeply emarginate at middle ( Fig. 9B View FIGURE 9 ); parameres as in Fig. 3P–Q................................... H View in CoL View FIGURE 3 . ulysses Burmeister

9´. Apex of the medial trough in the elevated section of the pronotum narrower, subequal in width to head in major males or narrower in minor males, weakly emarginate at middle ( Fig. 9A View FIGURE 9 ); parameres as in Fig. 3A–B..... H View FIGURE 3 . chevrolatii Burmeister