Ennearthron spenceri ( Hatch, 1962 )

Ennearthron spenceri ( Hatch, 1962)

Ennearthron victori Lopes-Andrade and Zacaro, 2003

Ennearthron melancholicum Rosa-Oliveira and Lopes-Andrade sp. n.

Incertae sedis The following seven names are here formally excluded from the genus and left as incertae sedis within Ennearthron abeillei Caillol ; Cis filum Abeille de Perrin ; Ennearthron hayashii Nobuchi ; Ennearthron Lohse ; Cis pruinosulus Perris ; Ennearthron reichei Abeille de Perrin ; and Ennearthron reitteri Flach Andrade and Zacaro 2003 ; Lawrence 2016).

These incertae sedis species were cited as doubtfully included in the genus by Lawrence (1971 noted that the Palearctic C. filum , E. hayashii and E. palmi are all quite small ( 1.4 mm or less), narrow cylindrical, with dentate protibial apices, the males with two small sex patches on the head; C. pruinosulus larger than the aforementioned three species and lacks dentate protibiae and cephalic sex patches in E. reitteri is short and broad, with a dual and indistinctly seriate elytral vestiture, a carinate prosternum lateral pronotal margins, and dentate protibial apices, and is likely unrelated to any of the aforementioned species.

Additionally, based on our translation of the original descriptions ( Abeille de Perrin 1874b; Caillol E. reichei , from Egypt, has the prosternum projecting in front of the coxae, protibiae slightly widened with a dentate outer apical angle and a spinose outer edge, and males possess one very long, directed cephalic horn and two broad, pointed pronotal horns, along with dense and distinct punctation contrast, E. abeillei , from Marseille ( France), is described as having very fine and dense punctation, and confused vestiture of short, golden, scale-like setae, and males without cephalic or pronotal tions. The description of E. reichei closely matches those of species in the Cis tricornis group, restricted to the American continent ( Borlini and Lopes-Andrade 2023), suggesting the possibility mislabelled specimens. Ennearthron abeillei was described as being similar in size and C. pruinosulus and may represent either a synonym or a closely related species.