Chimarrhodella spinosa, Holzenthal & Blanhik & Ríos-Touma, 2025

Chimarrhodella spinosa sp. nov.

Fig. 1 View Figure 1

Type material.

Holotype male: Ecuador: • Zamora Chinchipe: Tributary crossing Hwy E 682, N of Valladolid , Tapichalaca Reserve, 4.53371°S, 79.13041°W, el. 1898 m, 12.ii.2023, Ríos, Amigo, Huisman ( UMSP 000279961 View Materials ) ( MECN). GoogleMaps

Diagnosis.

Chimarrhodella spinosa is very closely related to C. costaricensis Blahnik & Holzenthal, 1992 , resembling it particularly in the structure of tergum X, which is membranous mesally and has sensillate lateral projections, each rounded apically, with the apex reflexed on its ventral margin to form a prominent, acute, ventrally projecting, spine-like projection at approximately midlength. Also, like C. costaricensis , and unlike other species in the galeata group of Chimarrhodella , it has the apices of the inferior appendage acute, resembling species of the peruviana group in this respect. Chimarrhodella spinosa is most readily distinguished from C. costaricensis by the spines of the phallic apparatus. Like C. costaricensis , it has a pair of short, symmetrically positioned, phallic spines basally, although slightly longer in length than in C. costaricensis ; it differs in having an additional pair of spinose tracts, composed of clustered, fine, needle-like spines, anterior to the paired phallic spines. These tracts of fine spines are a unique feature among known species of the genus. Chimarrhodella spinosa also differs from C. costaricensis in its somewhat larger size and longer inferior appendage, with their apicomesal margins more abruptly narrowed at approximately midlength.

Description.

Forewing length male 7.0 mm. Color pale stramineous, denuded (specimen in alcohol). Male. Segment IX, in lateral view, with anteroventral margin weakly produced, concavely excised mesally; posterolateral margins with weakly developed, setose, dorsolateral expansions at level of preanal appendage. Tergum X membranous mesally, shallowly notched at apex, laterally with pair of sclerotized, apically sensillate lobes; each lobe with large sclerotized, spine-like, anteroventrally directed projection. Preanal appendage elongate, but short in comparison to other species of Chimarrhodella ; inserted dorsad of widest point of posterolateral expansion of segment IX. Inferior appendage, in lateral view, elongate, linear; in ventral view, widest basally, narrowing apically, abruptly and concavely narrowed on mesal surface in apical half, apex with short, acute, mesally directed projection. Phallus with phallobase tubular, elongate, expanded basodorsally, extended posteroventrally; endotheca elongate, with well-developed, preapical, pleated, hood-like expansion; basally with pair of subequal, elongate, phallic spines, lacking basal enlargements, and additional paired tracts of fine, needle-like spines; phallotremal sclerite complex elongate, tube-like, weakly sclerotized overall, slightly more sclerotized basally; ventral margin with wide sclerotized strip, narrowing preapically, weakly forked and more sclerotized apically.

Female. Unknown.

Etymology.

The name refers to the tract of fine spines in the endotheca, a unique feature among known species of the genus.