Coenobita pseudorugosus Nakasone, 1988

Coenobita pseudorugosus Nakasone, 1988 View in CoL

( Figs. 3D‒F View Figure 3 , 4B View Figure 4 )

Habitat. Upper intertidal, sandy beach with gravel, dead coral, and leaf litter and vegetation.

Distribution. Cebu, Philippines. Indonesia: North

Halmahera, this is the first record in Indonesia.

Remarks. This is the first record of the species after its description by Nakasone in 1988. Among the species with stridulatory apparatus on the left cheliped, this species mostly resembles C. rugosus . The differences of these two species agree well with Nakasone’s (1988) observation: the lower margin of the palm of the left cheliped is straight in the distal half in C. pseudorugosus ( Fig. 3D View Figure 3 ), versus more convex in C. rugosus ( Fig. 3A View Figure 3 ); right male sexual tube longer and more slender than the left in C. pseudorugosus ( Fig. 3F View Figure 3 ), versus left and right male sexual tube similar, or right very slightly longer ( Fig. 3C View Figure 3 ). Furthermore, in C. rugosus the dark brown patch on the left cheliped palm is clearly indicated even after preservation ( Fig. 3A View Figure 3 ), while in C. pseudorugosus it is blurred ( Fig. 3D View Figure 3 ).