Gnathoclita (Gnathoclita) izerskyi, Gorochov, 2018

Gnathoclita (Gnathoclita) izerskyi View in CoL sp. nov.

( Figs 17–22 View Figs 17–29 , 30–34 View Figs 30–36 )

Etymology. This species is named after V. Izerskyy for his great help in organizing my work on the Peruvian Ensifera in 2017.

Type material. Holotype – male, PERU: Junin Department, Satipo Prov. , ~ 40 km NE of Satipo Town , environs of Calabaza Vill. , 11.507832°S, 74.811745°W, 2000–2200 m, on leaf of low bush in disturbed forest near river, at night, 1–4 December 2017, A. Gorochov, G. Irisov. Paratype – female, same data as for holotype.

Description. Male (holotype). General appearance most similar to that of G. (G.) peruviana Carl, 1921 . Head very high and not wide (its widest part located near bases of mandibles; Fig. 17 View Figs 17–29 ); upper rostral tubercle short, angular in profile and narrow (laterally compressed but with dorsomedian very narrow and rather deep concavity); lower rostral tubercle much lower, pressed to previous tubercle anteriorly and with almost flat anterior surface and narrowly rounded dorsal apex; ocelli moderately small (lateral ocelli shortly oval, distinct; median one almost round, less distinct); eyes moderately small, almost semiglobular; space between antennal cavities narrow (scape approximately 3.5 times as wide as this space); borders of these cavities almost as in previous taxa ( Rhammatopoda spp. ); epicranium under eyes and antennal cavities rather high, with large concavity on each gena and vertical keel-like convexity under each eye; clypeus wide and very high, with short upper portion lacking tubercles or processes on its dorsal part (or on epicranium near this part) and with high- er (longer) lower portion which gradually narrowing to clypeus; mandibles very long and arcuate, directed downwards/forwards, with proximal half having small rounded convexity near base (in G. peruviana , this convexity more conical and located closer to middle of mandible; for comparison see Figs 17, 18 and 23, 24 View Figs 17–29 ), and with distal parts (having mandibular teeth at apex) less strongly curved to each other than in G. peruviana . Pronotum almost indistinguishable from that of this species (see Figs 18, 19 and 25, 26 View Figs 17–29 ); tegmina reaching abdominal apex but distinctly narrower than in G. peruviana (in latter species, tegmina slightly protruding beyond abdominal apex) and with distinctly shorter stridulatory vein and clearly narrower mirrors (see Figs 19–21, 26–28 View Figs 17–29 , 30, 31, 35 View Figs 30–36 ); hind wings ( Fig. 30 View Figs 30–36 ) clearly not reaching abdominal apex (barely protruding beyond it in G. peruviana ); legs (including inner and outer tympana) almost indistinguishable from those of latter congener. Abdominal apex also similar to that of G. peruviana , but epiproct with roundly angular (not simply rounded) posterior part (see Figs 22 and 29 View Figs 17–29 ), each cercus with ventral hook less arcuate in posterior view and almost without concavity above this hook (see Figs 33 and 36 View Figs 30–36 ); genital plate with narrower and less notched apical part as well as with longer styles (see Figs 22 and 29 View Figs 17–29 ); genitalia completely membranous. Body colouration distinguished from that of G. peruviana by larger darkened parts of clypeus and pronotal disc as well as darker most part of tegminal membranes (see Figs 17–19 and 23–26 View Figs 17–29 ).

Female. Head with epicranium more or less similar to that of male but slightly narrower in lower part, with mouthparts distinctly shorter (less high) and almost unspecialized: clypeus clearly lower; mandibles almost not arcuate, directed downwards and without distinct convexities or tubercles. Pronotum, wings and legs more similar to those of male, but tegmina without stridulatory apparatus and insignificantly less narrow in middle part; abdominal apex with epiproct rounded almost as in male of G. peruviana but smaller, with cerci thinner and unspecialized, and with genital plate and ovipositor as in Figs 32, 34 View Figs 30–36 . Body colouration also similar to that of male, but tegminal membranes almost blackish.

Length (mm). Body: male 25.0, female 27.5; pronotum: male 4.5, female 5.2; tegmina: male 15.5, female 19.5; hind femora: male 14.3, female 16.7; ovipositor 12.5.

Comparison. The new species differs from the closely related G. peruviana in the shorter stridulatory vein and narrower mirrors in the male tegmina as well as some small characters of male mandibles, male abdominal apex and body colouration (see above). From all the other congeners, the new species is distinguished by: the tegmina more spotted, shorter and having narrower mirrors [from G. laevifrons Beier, 1960 ]; the head narrower, and tegmina longer [from G. sodalis Brunner-Wattenwyl, 1895 ]; by the anteroproximal mandibular tubercle distinctly smaller, and tegmina clearly shorter [from G. vorax (Stoll, 1813) ]; by the absence of tubercles or spines near the dorsal clypeal edge, and tegmina distinctly longer (from the subgenus Disceratus Scudder, 1869 ). It is necessary to note that in the electronic catalogue (OSF), this subgenus was again restored as a separated genus; it is a mistake, because some species of the subgenus Disceratus have intermediate characters between type species of this subgenus and representatives of Gnathoclita s. str. Morever, the too fragmented and confused classification proposed by Beyer (1960, 1963) seriously complicates the generic determination in this complex subfamily.