Guatteria friesiana (W.A.Rodrigues) Erkens & Maas

6. Guatteria friesiana (W.A.Rodrigues) Erkens & Maas View in CoL — Fig. 8a View Fig , 9 View Fig ; Plate 1c; Map 2 View Map 2

Guatteria friesiana (W.A.Rodrigues) Erkens & Maas (2008) View in CoL 404. — Guatteriopsis friesiana W.A. Rodrigues (1981) View in CoL 49, f. 1; Maas et al. (2007) 644. — Type: D. Coêlho INPA3609 (holo INPA), Brazil, Amazonas, Manaus, Igarapé do Passarinho, 14 Mar. 1956.

Tree 3–10 m tall, 4–10 cm diam; young twigs and petioles densely covered with a velutinous indument of erect, dark brown to blackish brown, long-persistent hairs up to 0.5 mm long. Leaves: petiole 2– 8 mm long, 2 – 3 mm diam; lamina narrowly elliptic to narrowly obovate, 12– 28 by 4 –9 cm (leaf index 2.7– 3.5), chartaceous to thinly coriaceous, very densely confluentverruculose, pale green to greyish green on both sides, glabrous above, sparsely covered with appressed brown hairs below, but rather densely to sparsely covered on primary vein, base cordate, apex acuminate (acumen 10–20 mm long), primary vein flat or almost so above, secondary veins indistinct, 10 –24 on either side, flat or slightly impressed above, indistinctly loop-forming over most of the length or the entire length, shortest distance between loops and margin 2 – 5 mm. Flowers in 1–2-flowered inflorescences in axils of leaves; pedicels 5 –10 mm long, 2 – 3 mm diam, densely covered with dark brown, appressed hairs, articulated at 0.3 –0.5 from the base, bracts c. 5, soon falling, the upper bract broadly to depressed ovate, c. 5 mm long; flower buds ovoid to conical, acute; sepals free, triangular to ovate-triangular, 10–16 mm long, outer side densely covered with dark brown, appressed hairs; petals cream in vivo, 125 127 narrowly ovate-triangular to narrowly oblong-ovate, outer petals 20– 25 by 8 –12 mm, inner ones 15–19 by 6– 8 mm, outer side densely covered with dark brown, appressed hairs; stamens c. 2 mm long, connective shield hairy. Monocarps 7–10, green, maturing red in vivo, ellipsoid, 19–23 mm long, sparsely covered with appressed and erect hairs, apiculate (apiculum c. 2 mm long), stipes 1–3 mm long. Seed ellipsoid, 13–19 by 6– 8 mm, dark, shiny brown, slightly sulcate at the base, slightly pointed at the apex.

Distribution — Brazil (Amazonas).

Habitat & Ecology — In non-inundated forest, on clayey to sandy soil. At elevations of 50–125 m. Flowering: February to May; fruiting: March to June.

Vernacular names — Brazil: Envireira ( Rodrigues & Loureiro 5908).

Note — Guatteria friesiana can be recognized by its leaves with a cordate base, obscure secondary veins, a velutinous indument on young twigs and shortly pedicellate flowers.