Penthicus iners ( Ménétriés, 1832 )

Penthicus iners ( Ménétriés, 1832)

( Figs 42–44 View FIGURE 42 View FIGURE 43 View FIGURE 44 )

Ménétriés 1832: 202 ( Heliophilus ); Gridelli 1954: 286 ( Lobodera ); Kühnelt 1957: 78 ( Lobodera ); Kaszab 1961: 116 ( Lobodera );

Medvedev 1964: 658 ( Lobodera iners iners ); Abdurakhmanov & Nabozhenko 2011: 109, 244, fig. 239; Nabozhenko et al.

2021: 437. = saginatus Ménétriés, 1832: 203 ( Opatrum ). = molestus Faldermann, 1836: 388. = obesus Kulzer, 1964: 223 , syn. n. ( Lobodera ); Kaszab 1969: 451 ( Lobodera ); Grimm 2015: 315. The complete list of references for the Caucasus up until 2010 is presented in Abdurakhmanov & Nabozhenko (2011).

Type material. Heliophilus iners ( now Penthicus iners ). Lectotype ( ZIN), sex unknown, designated here ( Figs 42A–C View FIGURE 42 ): “ heliophilus iners Menetr. Zouv. ” (handwritten by E. Ménétriés; now Zuvand, Azerbaijan), goldish circle, “ Lectotypus Helophilus iners Ménétriés 1832 des. M.V. Nabozhenko, 2024”.

Opatrum saginatum (now Penthicus saginatus ). Lectotype ( ZIN), sex unknown, designated here ( Figs 42D–E View FIGURE 42 ): “ Opatrum Saginatum Menetr. ” (handwritten by E. Ménétriés), goldish circle, “ Heliopates iners var. B”, “ Penthicus saginatus ”, “ L. iners Mén. Reiçhardt det.”, “ Lectotypus Opatrum saginatum Ménétriés 1832 des. M.V. Nabozhenko, 2024”. Ménétriés (1832) noted that he collected this species from Baku environs ( Azerbaijan). Abdurakhmanov & Nabozhenko (2011) indicated that P. iners was not found in Baku and environs after the record of E. Ménétriés and possibly extinct in this area.

Lobodera obesa ( now Penthicus obesus ). Holotype ( NHMB) ( Figs 42F, I View FIGURE 42 ): “11–62 3000 m Demavend ”, “ Persia Elburs-Gebg. Hüdepohl leg.”, “ Holotypus Lobodera obesa n. sp. det. H.Kulzer 1963”; 1 paratype ( HNHM) ( Figs 42G, H, J View FIGURE 42 ): “ Persia Elburs Gebg. Hüdepohl leg.”, “11–62 3000 m Demavend ”, “ Paratypus Lobodera obesa n. sp. det. H. Kulzer 1913”, “ Paratypus 1964 Lobodera obesa Kulzer ”.

Material (only from Iran). 1 specimen ( ZIN): Iran, Ardabil Prov., Sabalan Mt. [38°16′01″N, 47°50′13″E], iv.1874 (leg. G.F. Christoph) GoogleMaps ; 1 specimen ( ZIN): Iran, Semnan Prov., Tash N of Shahrud, 40 versts (= 42.6 km) north of Shahrud [now Tash-e Bala, 36°34ʹN, 54°40ʹE] (leg. G.F. Christoph) ; 4 specimens ( ZIN): Golestan Prov., ShahKuh Mt. (4000–10000 ft [= 1219–3048 m a.s.l.]) S of Astrabad (now Gorgan), vii (leg. G.F. Christoph) ; 1 specimen ( HNHM): “ Perses ”, “ Museum Paris , coll. M. Séllidot 1935” (determined as P. obesus ) ; 2 specimens ( HNHM): Iran, Tehran / Mazandaran provinces, Polour—Abali (between Polur and Abali, 35°50′55″N, 52°03′01″E – 35°46′11″N, 51°58′00″E), 2100–2800 m, 17.v.1970 (leg. Wittmer V. Bothmer) (determined as P. corpulentus ) GoogleMaps ; 2 specimens ( HNHM): Iran, East Azerbaijan Province, Kalan Arasbaran wildlife refuge (38°44′N, 46°42′E), 2450–2650 m, 6– 10.vi.1978 (leg. Mattenzone, Martens & Pieper) GoogleMaps ; 14 specimens ( HNHM): Iran, Alborz Mts, Demavend , 2500– 2300 m and 4000–4300 m, 22–25.viii.1966 (leg. V. Sbordoni) ; 3 specimens ( HMIM): Iran, Tehran Prov., Damavand , 23.vii.1970 (leg. Saf-Hasch) ; 6 specimens ( HMIM): Iran, East Azerbaijan Prov., Ahar city, Kalibar , 14.vi.1973 (leg. Rogabi) ; 1 specimen ( HMIM): Iran, Ardabil [erroneously as Azarbaijan on the label] Prov., Sabalan Mt. , H = 2800–3800 m, 25.vi.1973 (leg. Rogabi) ; 1♂, 1♀ ( PCMN): Iran , Zanjan Prov., 40 km NW Bonab, 36º47ʹ57ʺN, 48º43ʹ53.7ʺE, 9–10.v.2017 (leg. D.G. Kasatkin) GoogleMaps ; 2 ♀♀ ( PCMN) Iran, West Azerbaijan Prov., Urmia Lake env., near Rajan vill., 37°22ʹN, 44°51ʹE, 22.v.2017 (leg. D.G. Kasatkin) ; 4 specimens ( PCMN): Iran , Zanjan Prov., 36°45ʹ37ʺN, 48°48ʹ13ʺE, 9.v.2017 (leg. A. Zubov) GoogleMaps ; 7 specimens ( PCMN): Iran, East Azerbaijan Prov., Kandovan , 21.v.2017 (leg. A. Zubov) ; 1 specimen ( PCMN): Iran, Lorestan, Dorud , 25.v.2017 (A. Zubov) ; 9 specimens ( HMIM): Iran, Ardabil Prov., Alborz Mts., Susahab , Mirajin willage env., H= 2100–2300 m, 37°23ʹ13.3ʺN, 48°30ʹ28.0ʺE, 5.vii.2024 (leg. F. Eshraghi Mofrad) GoogleMaps .

Notes. Specimens, described as Lobodera obesa are conspecific to P. iners . Kulzer (1964) compared his species P. obesus with P. fartilis and P. persicus , but not with P. iners . Specimens of P. obesus have the pronotum widest at middle or slightly behind middle. The shape of the pronotum is variable in P. iners even in one population and the pronotum can be widest slightly ahead middle (lectotype of P. iners ), at middle or slightly behind middle. Thus, the following new synonymy is introduced: Penthicus iners ( Ménétriés 1832) = P. obesus ( Kulzer, 1964) , syn. n.

Distribution. Turkey (Hakkari Prov.) (PCMN), Azerbaijan (Talysh, Nakchivan), North Iran (from West Azerbaijan Province to Schahrud and Gorgan) ( Medvedev 1964, Abdurakhmanov & Nabozhenko 2011), Armenia ( Nabozhenko et al. 2021). The record of the species from Turkmenistan (Gaudan pass) ( Reichardt 1936) needs verification ( P. iners or similar P. turcomanicus G.S. Medvedev, 1964 , which was originally described as a subspecies of P. iners from Central Kopetdag ( Medvedev 1964)); we didn’t find specimens from Gaudan in ZIN collection. Kaszab (1969) listed two specimens of P. obesus from Dasht-e Arzhan (Fars Prov.), which are erroneously labeled. The species P. iners was listed as could potentially be found in China ( Ren & Yang 2006), but this is dubious information.