Veronica biloba L. var. glandulosissima Bornm.

Veronica biloba L. var. glandulosissima Bornm. View in CoL

Beihefte zum botanischen Centralblatt 22: 112 ( Bornmüller 1907a).

TYPE CITATION. — Iran, Kerman: “Mt. Kuh-i-Dschupar [= Kuh-i-Jupar, c. 29°55.8’N, 57°11.5’E], 7.V.1892 leg (Bornm. Exsicc. No. 5017; edit. 1894)”.

LECTOTYPE (designated here). — “ J. Bornmüller: Iter Persico-turcicum 1892-93 No. 5017 […] Persia, prov. Kerman: in monte Kuh-i-Dschupar. 28-3200 m.s.m. 1892. V. 7. Legit et determ.: J. Bornmüller ”, JE[ JE00000136 !*].

ISOLECTOTYPES (designated here). — BP[ BP348273 !], BR[ BR0000005422333 !*], HBG[ HBG512112 *], K[ K001070272 *], OXF!, P[ P04048967 *], PRC[ PRC453974 !*], STU!, WU!

Nomenclatural note. Veronica biloba is a polymorphic annual species occurring from Lebanon and southern Turkey to Mongolia. The indumentum is variably eglandular and/ or glandular and hexaploid and octoploid populations are known ( Albach 2008b). The closely related V. argute-serrata , occurring basically from the same region with the same variation in indumentum, is distinguished from V. biloba by dentate bracts. Based on this character, Fischer (1981) synonymized V. biloba var. glandulosissima with V. argute-serrata . However, this character is influenced by the environment. Own experiments demonstrated that seeds from classical V. biloba produce plants with dentate bracts when grown in nutrient-rich soil with sufficient water. Therefore, V. biloba and V. argute-serrata are likely to be synonymized after more detailed investigation and variants with divergent characters such as strong glandular indumentum are likely to be split.