Bolla ( Stolla ) madrea (R. Williams and E. Bell, 1940)

Bolla ( Stolla) madrea (R. Williams and E. Bell, 1940) View in CoL and Bolla ( Stolla) hazelae (Hayward, 1940) are species distinct from Bolla ( Stolla) zorilla (Plötz, 1886)

Genomic sequencing of the holotypes of Pholisora madrea R. Williams and E. Bell, 1940 (type locality in Peru: Madre de Dios, NVG-15097C09, GenBank barcode OR665731) and Pholisora hazelae Hayward, 1940 (type locality in Ecuador, NVG-18024G08, GenBank barcode OR665732) and a syntype of Antigonus zorilla Plötz, 1886 (type locality in Panama, NVG-18056H01, GenBank barcode OR665733), currently treated as conspecific in the genus Bolla Mabille, 1903 (type species Bolla pullata Mabille, 1903 , considered a junior subjective synonym of Staphylus imbras Godman and Salvin, 1896 ) reveals that while all three are in the same clade, they are genetically differentiated at the level not unprecedented even for subgenera ( Fig. 5 View Figure 5 ), e.g., COI barcode differences are: 8.2% (54 bp) for P. madrea and P. hazelae , 7.1% (47 bp) for P. madrea and A. zorilla , and 7.6% (50 bp) for P. hazelae and A. zorilla . Therefore, we propose that Bolla ( Stolla) madrea (R. Williams and E. Bell, 1940) , restored status, and Bolla ( Stolla) hazelae (Hayward, 1940) , restored status, are species distinct from Bolla ( Stolla) zorilla (Plötz, 1886) .