Tylodinus variabilis Luna-Cozar, 2014

12. Tylodinus variabilis Luna-Cozar , sp. nov.

( Figures 2 View FIGURES 1–11. 1–5 , 50–51 View FIGURES 44–51 , 107 View FIGURES 100–109 , 132 View FIGURES 132–133 )

Diagnosis. Length male, 3.4–4.0 mm, female, 3.2–4.0 mm. Width male, 1.6–1.7 mm, female 1.5–2.0 mm. As for T. buchanani with the follow exceptions: body clothed with short scales, light scales on vertex and pronotum frequently whitish, but in some specimens light brown. Head with overlapping scales on vertex; front moderately concave. Elytra with basal margin straight ( Figure 2 View FIGURES 1–11. 1–5 ), stria 10 complete, surface granulate, tubercles similar to T. buchanani . Metasternal tooth absent. Abdomen squamous, scales setigerous, ventrite 1 convex; ventrite 2 longer than 3–4 combined, convex. Hind femora length almost reaching elytral apex, front tibiae sinuate, middle and hind tibiae subparallel, premucro very small, hind tibiae with apical ¼ curved. Male genitalia ( Fig. 107 View FIGURES 100–109 ) in lateral view slender; in ventral view with lateral sides concave, subparallel; apical process in lateral view weakly curved. Female genitalia with tergite eight trapezoidal, apex slightly curved, sternite eight oblong, median membranous area subapical; spermatheca shallowly curved, ramus long and slender, nodulus slender.

Geographic distribution. México ( Chiapas).

Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 2300–2750 m.

Derivation of specific name. Name derived from the Latin word variabilis = variable.

Material examined. Total 95 males, 90 females. Holotype male ( CMNC): México: Chiapas, San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m ( 16°44'33''N, 92°41'38.4''W), 15.ix.1991, coll. R GoogleMaps . S. Anderson. Paratypes: México: Chiapas, Huixtán , 15 km E San Cristóbal, elev. 2500 m ( 16°44'49.6''N, 92°29'24.22''W), 28.v.2008, coll. LLAMA (Wa-A-05-1-24) GoogleMaps — 1 female ( CMNC); same locality ( 16.74717°N 92.49017°W), 28.v.2008, coll. LLAMA (Wa-A-05-1-48) GoogleMaps — 1 female ( CMNC); same locality ( 16.74698°N 92.49013°W), 28.v.2008, coll. LLAMA (Wa-A-05-1-16) GoogleMaps — 1 male ( CMNC); Bazom , elev. 2450 m ( 16°44'18.96''N, 92°29'18.24''W), 9.vii.2003, coll. R GoogleMaps . S. Anderson — 1 male ( CMNC); San Cristobal de las Casas Airport , elev. 2350 m ( 16°41'25.08''N, 92°31'48''W), 25.vii.2005, coll. R GoogleMaps . S. Anderson — 1 female ( CMNC); San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m ( 16°44'33''N, 92°41'38.4''W), 15.ix.1991, coll. R GoogleMaps . S. Anderson — 6 males, 6 females ( CMNC); same locality, 18.ix.1991, coll. R GoogleMaps . S. Anderson — 4 males, 2 females ( CMNC); Cerro Huitepec 5 km W San Cristóbal, elev. 2650 m ( 16°44'37''N, 92°41'23.3''W), 17.ix.1992, coll. R GoogleMaps . S. Anderson — 2 males, 5 females ( BMNH, CMNC); Corazón de María, elev. 2449 m ( 16°41'7.4''N, 92°32'11.4''W), 8.vi.2006, coll. J. L. Cozar — GoogleMaps 1 male ( ECOS); Huitepec , elev. 2466 m ( 16°44'54.2''N, 92°41'6.4''W), 5.v.2006, coll. J. L. Cozar — GoogleMaps 2 males, 2 females ( ECOS); same locality, 15.vi.2006, coll. J. L. Cozar — 1 male ( ECOS); same locality, 3.vii.2006, coll. J. L. Cozar — GoogleMaps 2 males ( ECOS); same locality ( 16.75181°N 92.68270°W), elev. 2480 m, 29.v.2008, coll. LLAMA (Wa-A-05-2-01) GoogleMaps — 2 males, 1 female ( CWOB); same locality ( 16.75104°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-18) GoogleMaps — 1 male, 1 female ( CMNC); same locality ( 16.75109°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-17) GoogleMaps — 1 female ( CMNC); same locality ( 16.75131°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-12) GoogleMaps — 1 female ( CMNC); same locality ( 16.75176°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-02) GoogleMaps — 3 males, 1 female ( CMNC); same locality ( 16.75163°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-05) GoogleMaps — 3 males, 5 females ( CMNC, ECOS); same locality ( 16.74689°N 92.48990°W), 29.v.2008, coll. LLAMA (Wa-A-05-1-01) GoogleMaps — 1 male ( CMNC); same locality ( 16.75018°N 92.68280°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-39) GoogleMaps — 2 males ( CMNC); same locality ( 16.75018°N 92.68284°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-40) GoogleMaps — 1 male ( CMNC); Huitepec Ocotal, Cerro La Cruz, elev. 2970 m ( 16°44'31.2''N, 92°41'37.7''W), 12.vi.2006, coll. J. L. Cozar — GoogleMaps 3 males, 2 females ( ECOS); Huitepec , 5 km W San Cristóbal, elev. 2750 m ( 16°44'33''N, 92°41'38.4''W), 14-25.ix.1992, coll. R GoogleMaps . S. Anderson — 2 females ( CMNC); same locality, 14.ix.1992, coll. R GoogleMaps . S. Anderson — 5 males, 2 females ( CMNC); Huitepec, S. Cristóbal , elev. 2480 m ( 16.75041°N 92.68270°W), 29.v.2001, coll. LLAMA (Wa-A-05-2-32) GoogleMaps — 1 male, 1 female ( CMNC); same locality ( 16.75032°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-34) GoogleMaps — 2 females ( ECOS); same locality ( 16.75045°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-31) GoogleMaps — 1 male, 2 females ( CMNC); same locality ( 16.75163°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-05) GoogleMaps — 1 female ( CMNC); same locality ( 16.75050°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-30) GoogleMaps — 1 male, 1 female ( ECOS); same locality ( 16.75032°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-34) GoogleMaps — 1 female ( CMNC); same locality ( 16.75077°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-24) GoogleMaps — 1 female ( CMNC); same locality ( 16.75018°N 92.68298°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-43) GoogleMaps — 1 female ( CMNC); same locality ( 16.75018°N 92.68303°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-44) GoogleMaps — 1 female ( CMNC); same locality ( 16.75018°N 92.68308°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-45) GoogleMaps — 1 male ( CWOB); same locality ( 16.75018°N 92.68317°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-47) GoogleMaps — 1 male ( CMNC); same locality ( 16.75018°N 92.68327°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-49) GoogleMaps — 1 female ( CMNC); same locality ( 16.75054°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-29) GoogleMaps — 1 male ( CMNC); same locality ( 16.75082°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-23) GoogleMaps — 1 male ( CMNC); Los Alcanfores, elev. 2450 m ( 16°44'12.47''N, 92°40'46.93''W), 25.iv.2006, coll. M. Girón — GoogleMaps 1 male ( ECOS); Reserva Huitepec , elev. 2400 m ( 16°44'12.5''N, 92°40'46.9''W), 18.xi.1993, coll. R GoogleMaps . Jones — 4 males, 1 female ( ECOS, UAQM); same locality, 18.xi.1993, coll. R GoogleMaps . Jones — 1 male ( ECOS); same locality, 16.ii.1994, coll. R GoogleMaps . Jones — 1 male ( ECOS); same locality, 16.ii.1994, coll. R GoogleMaps . Jones — 2 males, 2 females ( ECOS); same locality, 7.iv.1994, coll. R GoogleMaps . Jones — 1 females ( ECOS); same locality, 27.v.1994, coll. B. Gómez-G y O. L. Gómez-N— 2 males ( ECOS); same locality, 25.viii.1994, coll. B. Gómez-G— 2 males ( ECOS); same locality, 26.x.1994, coll. B. Gómez-G— 1 male, 1 female ( ECOS); same locality, 24.i.1995, coll. R GoogleMaps . Jones y B. Gómez-G— 1 female ( ECOS); same locality, 24.i.1995, coll. R GoogleMaps . Jones — 1 male ( ECOS); same locality, 24.i.1995, coll. R GoogleMaps . Jones-W y B. Gómez-G— 1 male ( ECOS); same locality, 27.iv.1995, coll. B. Gómez-G y M. Girón-I— 1 female ( ECOS); same locality, 14.vi.1995, coll. B. Gómez- G, M. Girón-I y A. Méndoza-R— 1 male ( ECOS); same locality, 27.vi.1995, coll. B. Gómez-G y M. Girón-I— 1 female ( ECOS); same locality, 5.vii.1995, coll. M. Girón — 1 male ( UAQM); same locality, 5.vii.1995, coll. R GoogleMaps . Jones — 1 male ( UAQM); same locality, 10.vii.1995, coll. R GoogleMaps . Jones — 1 female ( UAQM); same locality, 25.vii.1995, coll. R GoogleMaps . W. Jones — 4 males, 3 females ( ECOS, UAQM); same locality, 1.viii.1995, coll. A. Mendoza y M. Girón — 1 male ( UAQM); same locality, 15.viii.1995, coll. M. Girón — 1 female ( UAQM); same locality, 3.xi.1995, coll. B. Gómez y M. Girón — 1 male, 2 females ( UAQM); same locality, 14.xi.1995, coll. B. Gómez, M. Girón — 1 male, 1 female ( UAQM); same locality, 5.xii.1995, coll. B. Gómez y M. Girón — 1 male, 2 females ( UAQM); same locality, 6.ii.1996, coll. L. Ruiz-M y M. Girón-I— 3 males, 4 females ( ECOS, UAQM); same locality, 14.ii.1996, coll. L. Ruiz-M y M. Girón-I— 1 male ( ECOS); same locality, 14.ii.1996, coll. L. Ruiz-M y M. Girón-I— 2 males, 2 females ( UAQM); same locality, 7.vii.2003, coll. R GoogleMaps . S. Anderson — 1 female ( CMNC); same locality, 12.ii.2004, coll. J. L. Cozar, M. Girón — 1 female ( ECOS); same locality, 9.vi.2004, coll. J. L. Cozar, M. Girón — 1 male, 1 female ( ECOS); same locality, 11.vi.2004, coll. J. L. Cozar — 1 female ( ECOS); same locality, 2.vii.2004, coll. J. L. Cozar — 1 female ( ECOS); same locality, 12.vii.2004, coll. J. L. Cozar — 1 female ( ECOS); same locality, 13.vii.2004, coll. M. Girón — 1 male ( ECOS); same locality, 15.vii.2005, coll. R GoogleMaps . S. Anderson — 1 male ( CMNC); same locality, 9.vi.2006, coll. U. Caballero — 1 female ( ECOS); same locality, 12.vi.2006, coll. J. L. Cozar — 1 female ( ECOS); same locality, 17.vi.2006, coll. U. Caballero — 3 females ( ECOS); same locality, 11.vii.2007, coll. J. Longino — 1 female ( CMNC); same locality, 9.vii.2007, coll. R GoogleMaps . S. Anderson — 4 males, 4 females ( CMNC); same locality, 9.vii.2007, coll. M. G. Branstetter — 2 females, 1 male ( CMNC); same locality, 11.vii.2007, coll. R GoogleMaps . S. Anderson — 3 males, 2 females ( CMNC); Zoológico, elev. 2379 m ( 16°43'5.5''N, 92°42'9.7''W), 6.vi.2006, coll. J. L. Cozar — GoogleMaps 1 male ( ECOS); San Juan Chamula, Cerro Tzontehuitz, elev. 2800 m ( 16°49'10.2''N, 92°35'3.8''W), 15.vii.1994, coll. R GoogleMaps . Jones — 1 male ( UAQM) .