Tylodinus mutabilis Luna-Cozar, 2014

Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W. & León-Cortés, Jorge L., 2014, A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico, Zootaxa 3788 (1), pp. 1-63 : 38

publication ID	https://doi.org/10.11646/zootaxa.3788.1
publication LSID	lsid:zoobank.org:pub:7484832A-025B-41FA-9696-DF7531C01AA4
DOI	https://doi.org/10.5281/zenodo.5080176
persistent identifier	https://treatment.plazi.org/id/C27E87FF-FFBE-6129-FF14-DBAA20DEF8A9
treatment provided by	Felipe
scientific name	Tylodinus mutabilis Luna-Cozar
status	sp. nov.

Treatment

20. Tylodinus mutabilis Luna-Cozar , sp. nov.

( Figures 64–65 View FIGURES 60–67 , 113 View FIGURES 110–119 , 132 View FIGURES 132–133 )

Diagnosis. Length male, 2.3–2.8 mm, female, 2.4–2.7 mm. Width male, 1.1–1.4 mm, female 1.2–1.3 mm. Body approximately 2.3x longer than wide. Head with light brown to yellow scales; vertex with patch of overlapping whitish scales on middle of vertex. Pronotum in dorsal view feebly convex, outline with anterior and posterior constriction; disc somewhat hollowed. Elytra with basal margin sinuate; base of declivity with tubercle on I2 similar in size to tubercle on I4. Metasternum concave. Male abdomen with ventrite 1 concave, ventrite 2 with transverse line of scales along median line. Premucro absent. Male genitalia ( Figures 113 View FIGURES 110–119 ) with median lobe in lateral view weakly curved, moderately stout; apical process in lateral view moderately large (0.26–0.34 as long as median lobe), slightly curved, abruptly curved downward; in ventral view not constricted beyond the median orifice. Female genitalia with tergite eight trapezoidal, slightly curved at the apex, sternite eight trapezoidal; length of membranous median area 1/3 or ½ as long as basal plate; spermatheca very shallowly curved, ramus and nodulus indistinct.

Comments. This species is variable in the curvature of the front tibiae, sometimes being slightly sinuate; also, the external tooth on the middle tibia is weak or absent.

Geographic distribution. México ( Chiapas).

Habitat and elevation. Specimens were collected from leaf litter in various types of forest between 1100–1900 m.

Derivation of specific name. Name derived from Latin word mutabilis = changeable; inconstant.

Material examined. Total 23 males, 19 females. Holotype male ( CMNC): México: Chiapas, Villa Corzo, Ejido Sierra Morena, Reserva de la Biósfera La Sepultura , ( 16° 09' 10.6"N, 93° 35'25.1"W), elev. 1600 m, 19–20.vii.2005. R GoogleMaps . Anderson, oak pine ridge forest litter 2005-004A. Paratypes: México: Chiapas, same locality, elev. 1800 m, 18/ 19.vii.2007, R GoogleMaps . Anderson — 1 male, 1 female (( CMNC ); La Concordia, Reserva El Triunfo, Campamento el Quetzal , elev. 1850 m ( 15° 43'17"N, 92° 56'21.8"W), 17.v.2008, coll. R GoogleMaps . Anderson - 1 female ( CMNC); same locality, 19.vii.2007, coll. R GoogleMaps . Anderson — 1 female ( CMNC); same locality, elev. 1860m, 17.v.2008, coll. R GoogleMaps . S. Anderson — 1 male ( CMNC); same locality ( 15.72173N 92.94059W), elev. 1850m, 20.v.2008, coll. R GoogleMaps . S. Anderson — 1 female ( CMNC); 2 km SE Custepec , elev. 1520 m ( 15° 43'13.8"N, 92° 57'1.1"W), 17.v.2008, coll. LLAMA (Wa-A-02-1-13) GoogleMaps - 1 female ( ECOS); same locality ( 15.72078°N 92.95047°W), 17.v.2008, coll. LLAMA (Wa-A-02-1-18) GoogleMaps — 1 male ( ECOS); same locality ( 15.72098°N 92.95054°W), 17.v.2008, coll. LLAMA (Wa-A-02- 1-22) GoogleMaps — 1 male ( CMNC); same locality ( 15.72053°N 92.95100°W), 17.v.2008, coll. LLAMA (Wa-A-02-1-41) GoogleMaps — 1 female ( ECOS); same locality ( 15.72051°N 92.95104°W), 17.v.2008, coll. LLAMA (Wa-A-02-1-42) GoogleMaps — 1 male ( CMNC); same locality, elev. 1700 m ( 15.71522°N 92.93794°W), 17.v.2008, coll. LLAMA (Wa-A-02-2-39) GoogleMaps — 1 female ( CMNC); same locality, elev. 1700 m ( 15.71530°N 92.93820°W), 17.v.2008, coll. LLAMA (Wa-A-02-2- 40) GoogleMaps — 1 male ( CMNC); Villa Flores, Reserva La Sepultura, Ejido Sierra Morena , Parcela Escolar , elev. 1350 ( 16° 9'10.4"N, 93° 35'25.1"W), 23.vi.2006, coll. J. L. Cozar — 1 male ( CMNC); Villa Flores , Ejido Sierra Morena, R GoogleMaps . Biosfera La Sepultura , elev. 1400 m ( 16°9'10.4''N, 93°35'25.1''W), 17-18.vii.2003, coll. R GoogleMaps . S. Anderson — 1 female, 1 male ( CMNC); same locality ( 16°09.351’N 93°36.045’W), elev. 1450 m — GoogleMaps 1 female ( CMNC); same locality ( 16°09.856’N 93°36.291’W), elev. 1497 m — GoogleMaps 3 males, 1 female ( CMNC); same locality ( 16.16121°N 93.60024°W), elev. 1137 m — 2 males, 1 female ( CMNC, ECOS); same locality, elev. 1235 m, 19-20.vii.2005, coll. R GoogleMaps . S. Anderson — 1 male, 1 female ( CMNC); same locality, elev. 1350 m — 1 male, 2 females ( CMNC); same locality, elev. 1450 m — 2 males, 1 female ( CMNC); same locality, elev. 1600 m — 1 male, 1 female ( BMNH, CMNC); same locality ( 16.15976°N 93.60505°W), 12.v.2008, coll. LLAMA (Wa-A-01-1-26) GoogleMaps — 1 male ( ECOS); same locality ( 16.16044°N 93.60516°W), 12.v.2008, coll. LLAMA (Wa-A-01-1-44) GoogleMaps — 1 male ( CMNC); same locality ( 16.15458°N 93.60080°W), 12.v.2008, coll. LLAMA (Wa-A-01-2-05) GoogleMaps — 1 male ( ECOS); same locality ( 16.15276°N 93.60079°W), 12.v.2008, coll. LLAMA (Wa-A-01-2-36) GoogleMaps — 1 male ( CMNC); same locality ( 16.15287°N 93.60059°W), 12.v.2008, coll. LLAMA (Wa-A-01-2-41) GoogleMaps — 1 female ( ECOS); same locality ( 16.15292°N 93.60051°W), 12.v.2008, coll. LLAMA (Wa-A-01-2-43) GoogleMaps — 1 female ( CMNC); Sierra Morena , elev. 1550 m (16.1529°0'N, 93.5903°0'0''W), 17-18.vii.2003, coll. R . S. Anderson — 1 female ( CMNC) .