Rhyacophila retracta Martynov 1914

Rhyacophila retracta Martynov 1914 View in CoL

( Figs 8A–8C View FIGURES 8–11 , 12A–12D View FIGURES 12–15 , 16A View FIGURE 16 )

Rhyacophila retracta Martynov 1914 View in CoL , 75–77, male, Russia ( Altai); Ross 1956, 76, 103, 121, male; Botosaneanu 1970, 276– 277, 285, 328, female; Lepneva 1964, 250–255, larva; Schmid 1970, 90, 132, pl. 41, male; Tani, 1977, 198, male; Emoto 1979, 559–562, male, female; Tanida 1985, 181, 183, larva; Arefina 1997, 24, 31, male, female; Arefina 2001, 33, 44, female; Hattori 2005, 417, 422, 431, male, female, larva; Nozaki 2016, 311, 313, male; Kang 2020, 218–219, male, larva.

Rhyacophila uenoi Tsuda 1940 , 123–124, male, Hokkaido (Kamikawa). Synonymized by Ross (1956).

Rhyacophila uchidai Kobayashi 1989 View in CoL , 4–5, male, Korea ( Gyeongsangnam-do). Synonymized by Nozaki et al. (2019).

Diagnosis. Rhyacophila retracta Martynov 1914 is clearly distinguished from other species of the R. retracta Species Group by the following unique genital characters: In males, the short, trilobed apicodorsal lobe of segment IX ( Figs 8A, 8B View FIGURES 8–11 ); in females, segment VIII with a deep dorsal cleft and a pair of shallow lateral clefts ( Figs 12A, 12B View FIGURES 12–15 ).

Specimens examined. Hokkaido: 1 female, Horokaishikari-gawa, Kamikawa-cho , 6.vii.1993, M. Sato ( SPMN) : 1 female, Yamabe-nishi, Furano-shi , 16.vii.1996, T . Kishimoto ( SPMN) ; 1 female, Shikaribetsu, Shikaoicho , 27.viii.1995, unknown collector ( SPMN) ; 1 male, Shirogane-no-taki, Ofuyu, Hamamasu-ku, Ishikari-shi , alt. 5 m, 31.vii.2006, T . Ito & T . Hattori ( SPMN) ; 1 male, 4 females, Ponbetsu-gawa, Ikushunbetsu, Mikasa-shi , 22.vi.1997, T . Ito & A. Ohkawa ( SPMN) ; 1 male, 1 female, Takino, Sapporo-shi , 28.vii.1978, T . Ito ( SPMN) ; 1 male, Jozankei, Sapporo-shi , 23.viii.1995, H. Nishimoto ( SPMN) ; 2 males, 6 females, Ôtaki, Date-shi , 24.viii.1995, H. Nishimoto ( SPMN) ; 32 males, 16 females, Ken-ichi-gawa, Kumaishi-ayukawa-cho, Yakumo-cho , alt. 50 m, 23.vi.1997, T . Ito & A. Ohkawa ( SPMN) ; 3 males, same locality, 22.vii.1997, same collectors ( SPMN) ; 1 male, same locality, 25.viii.1997, same collectors ( SPMN) ; 1 male, Himekawa, Mori-machi , 29.ix.1993, T . Ito ( SPMN) ; 4 males, 9 females, same locality, 24.vi.1997, T . Ito & A. Ohkawa ( SPMN) ; 1 male, Shamisen-daki, Shikabe-cho , 11.ix.2006, T . Ito ( SPMN) ; 1 female, Uzurakawa, Assabu-cho , 30.v.2003, T . Ito & A. Ohkawa ( SPMN) ; 1 female, Kouzurakawa, Assabu-cho , 22.vi.2003, T . Ito & A. Ohkawa ( SPMN) ; 2 females, Ikusagawa, Nanae-cho , 23.vi.2003, T . Ito & A. Ohkawa ( SPMN) ; 1 male, 10 female, same locality, 11.ix.2006, T . Ito ( SPMN) ; 1 female, Idesu-kawa, Siriuchi-cho , 8.vii.1978, T . Ito ( SPMN) ; 1 male, 1 female, Mt. Horoshiri, Toyonuka, Biratori-cho , 25.vii.2017, K. Inazu (NKa). Tsushima Islands : 1 male, Azu-kawa, Izuhara-machi, Tsushima-shi , 3.vi.1984, M. Kobayashi ( KPM-NKMK77850 ) .

Distribution. Rhyacophila retracta is broadly distributed in Hokkaido, Tsushima Islands, Korea, northeastern parts of China, Sakhalin, Kuril Islands, continental parts of Russian Far East and southern Siberia ( Fig.16A View FIGURE 16 : Yang et al. 2005; Ivanov 2011; Taira et al. 2025). Kobayashi (1985) recorded females as this species from Kanagawa, central Honshu, but I identified them (KPM-NKMK76740) deposited in the Kobayashi collection of KPM as an unknown species belonging to the Rhyacophila angulata Species group Ross 1956.

Japanese name. Ueno-nagare-tobikera.