Tipula ( Vestiplex ) balioptera Loew, 1863

Tipula ( Vestiplex) balioptera Loew, 1863 View in CoL

Tipula balioptera Loew, 1863 View in CoL - Loew 1863: 284; Alexander 1915: 460;

Tipula ( Vestiplex) balioptera View in CoL : Crampton et al. 1942, Alexander 1965, Starkevich and Paramonov 2016, Starkevich et al. 2020, Brodo et al. 2023, Kolcsár et al. 2023;

= Tipula ( Vestiplex) laccata laccata View in CoL : Savchenko 1964 nec Lundstrom and Frey 1916 (partim), after Starkevich and Paramonov (2016);

= Tipula ( Vestiplex) tchukchi View in CoL : Mannheims 1971 nec Alexander 1934 (partim), after Starkevich and Paramonov (2016). According to Mannheims (1971), T. ( V.) tchukchi View in CoL = T. ( V.) laccata View in CoL : Lackschewitz 1936 (partim); Mannheims 1953 (partim); Savchenko 1964 (partim); nec Lundstrom and Frey 1916;

= Tipula ( Vestiplex) tchukchi View in CoL : Salmela 2008, Salmela 2011, Salmela 2012, Salmela and Petrašiūnas 2014;

Tipula ( Vestiplex) bo Mannheims, 1967 View in CoL - Mannheims 1967: 148 syn. nov. stat. res. Synonymy with T. ( V.) tchukchi Alexander, 1934 View in CoL after Mannheims and Savchenko (1973) = laccata Lackschewitz 1936 View in CoL (partim), Mannheims 1953 (partim), Savchenko 1964 (partim), nec Lundstrom and Frey 1916;

Tipula perretti Alexander, 1928 View in CoL - Alexander 1928: 98 syn. nov.; Tipula ( Vestiplex) perretti View in CoL : Crampton et al. 1942.

Materials

Type status: Lectotype. Occurrence: recordedBy: Kennicot; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Canada; verbatimLocality: English River; Record Level: institutionCode: MCZ

Type status: Other material. Occurrence: recordedBy: I. Kakko; individualCount: 2; sex: male, female; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Bombovarri SE; Event: verbatimEventDate: 15 VII 2010; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU391

Type status: Other material. Occurrence: recordedBy: I. Kakko; individualCount: 2; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Bombovarri SE; Event: verbatimEventDate: 15 VII 2010; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU392

Type status: Other material. Occurrence: recordedBy: A. Haarto; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Urtasvaara; Event: verbatimEventDate: 15 VII 2009; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU408

Type status: Other material. Occurrence: recordedBy: J. Salmela; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Tomuttioja; Event: verbatimEventDate: 11 VI – 19 VII 2009; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU824

Type status: Other material. Occurrence: recordedBy: J. Salmela; individualCount: 1; sex: female; associatedSequences: BOLD sample ID JES- 20110086; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Pahtavaara; Event: verbatimEventDate: 11 VI – 19 VII 2009; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU1858

Type status: Other material. Occurrence: recordedBy: J. Salmela; individualCount: 1; sex: male; associatedSequences: BOLD sample ID JES- 20110129; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Enontekiö, Tomuttioja; Event: verbatimEventDate: 11 VI – 19 VII 2009; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU1896

Type status: Other material. Occurrence: recordedBy: J. Salmela; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Utsjoki, Galddasjohka; Event: verbatimEventDate: 18 VI – 19 VII 2007; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU825

Type status: Other material. Occurrence: recordedBy: A. Rinne; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Finland; verbatimLocality: Utsjoki, Teno; Event: verbatimEventDate: 22 VI – 3 VII 2006; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.TIPU851

Type status: Other material. Occurrence: recordedBy: A. Järvinen; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Troms, Storfjord, Skibotn, Veajetnibba S; Event: verbatimEventDate: 24 VI – 16 VII 2017; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.ins2018-168

Type status: Other material. Occurrence: recordedBy: A. Järvinen; individualCount: 1; sex: male; occurrenceID: 8016767E-E022-557C-BF0F-F718132F850F; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Veajetnibba S; Event: verbatimEventDate: 19 VII – 12 VIII 2017; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.ins2018-199

Type status: Other material. Occurrence: recordedBy: A. Järvinen; individualCount: 3; sex: 2 males, 1 female; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Veajetnibba S; Event: verbatimEventDate: 16 VII – 19 VII 2017; Record Level: institutionCode: LMM; source: http://tun.fi/NVO.ins2018-205

Type status: Other material. Occurrence: catalogNumber: 721756 ; recordedBy: K. M. Olsen; individualCount: 1; sex: male; occurrenceID: F7C421F7-3612-5A86-8672-1B18AE8B97C9; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Troms, Kåfjord, N Nuorrtit Gussačohkka; Event: verbatimEventDate: 12 VII 2022; Record Level: institutionCode: PCKMO

Type status: Other material. Occurrence: catalogNumber: 718592 ; recordedBy: K. M. Olsen; individualCount: 5; sex: 3 males, 2 females; occurrenceID: 6797498A-FFE2-5196-A7AB-00AA10C38760; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: N Nuorrtit Gussačohkka; Event: verbatimEventDate: 1 VII – 12 VII 2022; Record Level: institutionCode: PCKMO

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18002 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Jotkajavre; Event: verbatimEventDate: 3 VII 1924; Record Level: institutionCode: TROM

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18001 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Jotkajavre; Event: verbatimEventDate: 9 VII 1924; Record Level: institutionCode: TROM

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18004 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Jotkajavre; Event: verbatimEventDate: 9 VII 1924; Record Level: institutionCode: TROM

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18006 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: female; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Jotkajavre; Event: verbatimEventDate: 9 VII 1924; Record Level: institutionCode: TROM

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18003 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Jotkajavre; Event: verbatimEventDate: 15 VII 1924; Record Level: institutionCode: TROM

Type status: Other material. Occurrence: catalogNumber: TROM TSZD 18005 ; recordedBy: T. Soot-Ryen; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) balioptera Loew, 1863 ; Location: country: Norway; verbatimLocality: Finnmark, Alta, Bojobæski; Event: verbatimEventDate: 16 VII 1924; Record Level: institutionCode: TROM

Type status: Holotype. Occurrence: catalogNumber: No. 2559 ; individualCount: 1; sex: male; preparations: Antenna and wing on slide mounted, USNM; Taxon: scientificName: Tipula ( Vestiplex) perretti Alexander, 1928 ; Location: country: Canada; verbatimLocality: Hopedale, Labr; Record Level: institutionCode: CNC

Type status: Paratype. Occurrence: catalogNumber: Type # 6365: 2; MZLU- 110421 ; recordedBy: A & S Ulfstrand; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) bo Mannheims, 1967; Location: country: Sweden; verbatimLocality: Lycksele Lappmark, Vindelälven, N Tjatsemvare, 45 km NW Ammarnäs; Event: verbatimEventDate: 23 VII 1965; Record Level: institutionCode: MZLU; source: https://www.flickr.com/photos/tags/mzlutype06365/

Type status: Paratype. Occurrence: catalogNumber: Type # 6365: 3; MZLU- 110422 ; recordedBy: A & S Ulfstrand; individualCount: 1; sex: male; Taxon: scientificName: Tipula ( Vestiplex) bo Mannheims, 1967; Location: country: Sweden; verbatimLocality: Lycksele Lappmark, Vindelälven, N Tjatsemvare, 45 km NW Ammarnäs; Event: verbatimEventDate: 23 VII 1965; Record Level: institutionCode: MZLU; source: https://www.flickr.com/photos/tags/mzlutype06365/

Type status: Paratype. Occurrence: catalogNumber: Type # 6365: 4; MZLU- 110423 ; recordedBy: A & S Ulfstrand; individualCount: 1; sex: female; Taxon: scientificName: Tipula ( Vestiplex) bo Mannheims, 1967; Location: country: Sweden; verbatimLocality: Lycksele Lappmark, Vindelälven, N Tjatsemvare, 45 km NW Ammarnäs; Event: verbatimEventDate: 23 VII 1965; Record Level: institutionCode: MZLU; source: https://www.flickr.com/photos/tags/mzlutype06365/

Taxon discussion

The first record of T. ( V.) balioptera out of the Nearctic Region was published by Starkevich and Paramonov (2016) with taxonomic account as follows:

= Tipula ( Vestiplex) laccata laccata : Savchenko 1964 nec Lundstrom and Frey 1916 (partim), after Starkevich and Paramonov (2016).

A single male specimen from Yakutyia, Russia was detected amongst material identified as T. ( V.) laccata by E. N. Savchenko ( Savchenko 1964). Another partim of the T. ( V.) laccata material belonged to T. ( V.) aldrichiana , a species which was also recorded in the Palaearctic for the first time ( Starkevich and Paramonov 2016). Later, T. ( V.) balioptera was recorded in Mongolia ( Starkevich et al. 2020) and Norway ( Kolcsár et al. 2023).

Following Mannheims (1971), all material published by Savchenko (1964) as T. ( V.) laccata was synonymised with T. ( V.) tchukchi Alexander, 1934 , including some other material listed below:

Tipula ( Vestiplex) tchukchi Alexander, 1934

= laccata Lackschewitz 1936 (partim), Mannheims 1953 (partim), Savchenko 1964, nec Lundstrom and Frey 1916, after Mannheims (1971).

Finally, another species T. ( V.) bo Mannheims, 1967 was synonymised by Mannheims and Savchenko (1973) with the taxonomic list as follows:

Tipula ( Vestiplex) tchukchi Alexander, 1934

= laccata Lackschewitz 1936 , Mannheims 1953, Savchenko 1964, nec Lundstrom and Frey 1916, after Mannheims and Savchenko (1973).

As can be seen from the taxonomic records ( Mannheims 1971, Mannheims and Savchenko 1973), besides T. ( V.) bo, three partims of T. ( V.) laccata material were listed as T. ( V.) tchukchi .

After examination of type material of T. ( V.) bo and based on morphology analysis, we found that this species belongs to T. ( V.) balioptera and thus is a junior synonym:

Tipula ( Vestiplex) bo Mannheims, 1967 View in CoL - Mannheims 1967: 148 syn. nov. stat. res.

= laccata Lackschewitz 1936 (partim), Mannheims 1953 (partim), Savchenko 1964 (partim), nec Lundstrom and Frey 1916.

Subsequently, T. ( V.) tchukchi is exluded from the list of fauna from Norway, Sweden and Finland.