Strongylium (s. str.) suwawa, Telnov & Masumoto, 2024

Strongylium (s. str.) suwawa sp. nov. ( Figs 6 View Fig , 9D–F View Fig )

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Type material designated. Holotype ♂ BMNH: INDONESIA: SULAWESI UTARA, Dumoga – Bone N.P. March 1985. [printed] // ‘Clarke’ Camp Lower montane forest, 1140 m. [printed] // R.Ent.Soc.Lond. PROJECT WALLACE B.M. 1985–10 [printed] // 013663806 [printed, supplemented with QR code]. The right antenna is completely missing.

Paratypes 2 specimens. 1♂ BMNH: same labels as holotype but 013663804; 1 BMNH [not sexed, presumably a female]: INDONESIA: SULAWESI UTARA, Dumoga – Bone N.P. March 1985. [printed] // ‘Clarke’ Camp Lower montane forest, 1140 m. [printed] // At light [printed] // R.Ent.Soc.Lond. PROJECT WALLACE B.M. 1985–10 [printed] // 123 51 [printed, label pink] // not Bradymerus [handwritten, in part underlined] det. SCHAWALLER 2008 [printed] //? Hexarhopalus n. sp. [handwritten] det. SCHAWALLER 2009 [printed]; 013663805 [printed, supplemented with QR code] .

Derivatio nominis. Named after Suwawa, the Philippine language spoken in North Sulawesi in Bogani Nani Wartabone (Dumoga Bone) National Park, where the new species was found. Noun in apposition.

Measurements. Holotype male, total body length 11.8 mm; head length 1.5 mm, maximum head width across compound eyes 1.9 mm, pronotal length 2.1 mm, maximum pronotal width 4 mm, elytral length 8.2 mm, maximum combined width 4.4 mm. Paratypes 10.8–12.2 mm long.

Description. Holotype male. Forebody uniformly black-brown dorsally and ventrally. Elytra and abdomen brown. Head trapezoid, transverse, subopaque dorsally and ventrally. Labrum truncate at anterior margin. Epistoma truncate at anterior margin. Moderately deep, concave impression at place of frontoclypeal suture. Antennal insertion concealed in dorsal view beneath raised gena. Compound eye large, anterior margin broadly emarginate at antennal insertion and genal canthus, subtruncate at posterior margin, strongly protruding from lateral, moderately - from dorsal outline of head. Minimum interocular distance about 2.1–2.2× as wide as length of dorsal eye portion. Tempus short, subparallel, about as long as shortest eye length. Head dorsum densely punctate with irregularly shaped, moderately deep punctures. Intervening spaces subopaque, smooth, in par elevated, much narrower than punctures. Head dorsal setation dirty whitish, short and strongly C-like curved posteriorly, suberect, setae slightly clavate apically. Antenna moniliform, slightly widened distally, exceeds level of mesocoxa when directed posteriorly. Basal antennomere about 3–3.3× as long as antennomere two. Antennomere two small, slightly transverse. Antennomere three 4.7× as long as antennomere two, 1.3–1.4× as long as antennomere four. Antennomeres 3– 6 slender, antennomere 7 widened distally, antennomeres 8–10 widened, 7–11 with sensorial fields of punctures covered with dense short whitish setae ( Fig. 6D View Fig ). Terminal antennomere cylindrical, rounded apically, about 0.8× as long as penultimate antennomere. Terminal maxillary palpomere

21 small, strongly triangular. Pronotum strongly transverse, opaque dorsally and laterally, widest across midlength, converging anteriad and posteriad (nearly subparallel in front of posterolateral angles), broadly emarginate at anterior, sinuous at posterior margin, strongly convex in dorsal aspect but flattened in front of base. Anterior pronotal edge not margined, somewhat raised-deflected medially, tuberculate. Posterior pronotal edge not margined, irregularly punctate, somewhat raised at some places. Anterolateral angles obtuse angulate, posterolateral angles nearly right-angled in dorsal view, tip rounded. Lateral pronotal edge poorly defined as a carina, not margined, not visible in dorsal view, somewhat obtuse-angulate medially in dorsal view, in lateral view only distinct in basal half of pronotum, irregular and interrupted in anterior half. Moderately deep median longitudinal impression on pronotal disc except at base, disc bulged in dorsal aspect both sides of impression, shallowly impressed beyond each bulge near base. Pronotal punctures irregularly shaped, large, shallow. Intervening spaces opaque, raised (corrugate), much narrower than punctures except at sparser punctured median impression area. Bulges of pronotal disc particularly dense punctate-corrugate, with glossy and glabrous irregularly shaped and sized drop-like tubercles ( Fig. 6C View Fig ). Dorsal pronotal setae as those on head, seta does not surpass length of corresponding puncture. Scutellar shield small, narrow, rounded at posterior margin, opaque, minutely punctate and densely setose except for narrowly subglabrous midline. Elytra widest across postmedium, moderately glossy, subparallel, dorsally convex. Elytral surface with deepened, punctured longitudinal striae (eight complete to nearly complete and one short scutellar stria visible in dorsal view) and slightly elevated interstriae. Striae 4–5 shortest and conjoinning in apical fourth. Punctures in striae very deep, irregularly quatrefoil-shaped ( Fig. 6E View Fig ). Interstriae slightly elevated, opaque

22 microcorrugate, with numerous small drop-like glossy tubercles ( Fig. 6E View Fig ). Elytral lateral margin not visible in dorsal view. Elytral setae dirty yellowish, sparse, apically slightly clavate, C-like curved posteriorly, suberect, also present in striae. Epipleuron complete, moderately wide, opaque, corrugate and with sparse dirty yellowish appressed setae. Metathoracic wings fully developed (functional). Meso-, metathorax and abdomen opaque. Abdominal sternites densely punctate, delicately setose. Legs very long and slender. Femur slender, not clavate, densely punctate. Tibia shorter than corresponding femur, nearly straight, not widens distally. Metatibia shallowly on inner basal edge with patch of short yellowish setae

( Fig. 6F View Fig ). Tarsus long, shorter than corresponding tibia, underside with moderately long yellowish setae. Terminal tarsomere of pro- and mesothoracic leg about as long as combined length of remaining corresponding tarsomeres. Male sternite VII broadly rounded at posterior margin. Aedeagus as in fig. 9D–F.

Sexual dimorphism. Unknown.

Intraspecific variability. The setose patch at inner basal edge of metataibia is not present (worn?) in one of the paratypes, but the setae are significantly longer in another paratype ( Fig. 6G View Fig ) than in the holotype.

Differential diagnosis. This new species is unique among other congeners but strongly resembles S. (s. str.) napolovi sp. nov. (described above). The main distinguishing features of S. suwawa sp. nov. are the elytral setae not arranged into longitudinal rows on the elytral interstriae, the pronotum with glossy and glabrous tubercles on its highest portions, the interocular distance much wider (2.1–2.2× as wide as length of dorsal eye portion versus about 1.3× in S. napolovi sp. nov.), the comparatively longer antenna, the less strongly elevated elytral interstriae and the denser drop-like tubercles on them.

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Distribution. Sulawesi (central part): North Sulawesi Province.

ACKNOWLEDGEMENTS

We are indebted to Matthias Hartmann ( NME) for providing the highly interesting material described and discussed in the present paper for our study and for the donation of some paratypes. We are grateful to the editor and anonymous reviewers for valuable comments which helped to improve the overall quality of the manuscript .