Hygromia (Riedelia) limbata limbata ( Draparnaud, 1805 )

Hygromia (Riedelia) limbata limbata ( Draparnaud, 1805) View in CoL

( Figs 5–6; Supporting Information, Fig. S6)

Helio limbata Draparnaud, 1805: 100 . Type locality: ‘dans l’Agénois, et dans le Sorézois’.

Type material: One of the three syntypes ( NHMW /MO/14821) was selected and designated as a lectotype. Tis is most probably the specimen illustrated by Draparnaud (1805: plate VI, fig. 29), having a characteristic white band, also visible inside the aperture ( Fig. 5B). Two remaining specimens become paralectotypes ( Fig. 5D–E). Te type locality cannot be specified further (see labels Fig. 5B) .

Helio odeca Bourguignat, 1882: 69 , 314–315. Type locality: ‘Hendaye et Bayonne, dans les Basses-Pyrénées; vallée du Clain, près de Poitiers, dans la Vienne (col. Bourguignat)’.

Type material: Te well-preserved specimen from Hendaye was selected and designated as a lectotype MHNG-MOLL-116286 ( Fig. 6A). Te second specimen is the paralectotype. Te type locality is restricted to Hendaye.

? Helio veprium Bourguignat, 1882: 67 , 314. Type locality: ‘Eaux-Bonnes, dans les Basses Pyrénées; Ascros, au-dessus de Roquesteron, dans les Alpes-Maritimes (col. Bourguignat)’. Type material: It is likely that this nominal taxon includes several species because Bourguignat in Locard (1882) based it on specimens from two different geographical regions. In one of them, Alpes-Maritimes, Riedelia species do not occur. Terefore, in order to fix the application of the name, the preserved syntype, originating from Eaux-Bonnes, is designated here as a lectotype MHNG-MOLL-116240 ( Fig. 8A) and the type locality is restricted to Eaux-Bonnes.

Description

Shell width 10.9–17.7 mm, shell height 7.8–12.8 mm, whorls 5.2–6.6. Other measurements are presented in Table 1. Shell shape sub-globose with a varying height of spire, may be translucent. Last whorl slightly keeled, ofen with a light spiral band at the periphery. Aperture slightly reflected with a whitish or pinkish lip. Umbilicus usually partly (65.5% of cases) or completely (29.9%) covered by a reflected columellar margin; if open very narrow. Sculpture unevenly developed: ridges less visible on the lower side than on the upper side, where they have a different density ( Fig. 14). Love dart long, about 4–5 mm, arched ( Fig. 21C–F). Shell and genitalia features are presented in Table 2.

Distribution

Tis study verified the occurrence of the species in northern Spain, and southern, south-western, and northern France, including 42 populations in total (Supporting Information, Table S1; Fig. 1).

Habitat

According to Welter-Schultes (2012), habitats include forests, shrubs, and hedges in lower vegetation. Also found in moist, grassy habitats among herbage and ground liter in gardens and other disturbed places. Likewise, our samples were from such habitats, usually from semi-open, but also open and shaded places. Twenty-nine populations were found in natural or seminatural habitats and 11 in artificial environments, at altitudes from sea level to more than 1500 m a.s.l.