Mimogonellus dreybrodti, Steiner, 2017

Mimogonellus dreybrodti View in CoL , new species ( Figs 1–2 View FIGURE 1 View FIGURE 2 )

Type material. Holotype: LAOS: ♂: ‘111/09-, 13.01.2009, N20°14'56.2", E103°23'47", ( Tham Nam Lot, S. Ban Boumfart Village ), Vieng Thong, H (o)uaphan(h) Prov. GoogleMaps , Laos, Col : H. Steiner, Northern Laos-European Cave Project 2009’ ( SNUC) . Paratypes: LAOS: 2 ♂♂, same collecting data as the holotype ( SNUC) .

Comparative diagnosis. Mimogonellus dreybrodti can be readily separated from the two Asian congeners, i.e., M. japonicus Naomi , and M. yasutoshii Naomi , by the much larger body size ( 4.96–5.13 mm vs. 3.0– 3.2 mm), more strongly curved aedeagal median lobe, and much longer parameres each exceeding the apex of the median lobe.

Description. Male. Body ( Fig. 1A View FIGURE 1 ) elongate, cylindrical; length 4.96–5.13 mm; forebody length 2.40–2.42 mm; reddish brown to dark reddish brown, tarsi and mouthparts lighter in color.

Head ( Fig. 1B View FIGURE 1 ) longer than wide, length 0.76–0.81 mm, width 0.72–0.74 mm; dorsum sparsely covered with large and shallow punctures, with long and yellow setae directed posteriorly, surface of disc between punctures polished and lacking microsculpture; mediobasal portion, area posterior to eyes, genae, and gular area with dense reticulate microsculpture; gular suture divergent for short distance at base, fused anteriorly; eyes slightly prominent, reniform in lateral view, each eye composed of about 130 ommatidia. Antennae ( Fig. 1C View FIGURE 1 ) reaching 1/3 elytral length when stretched posteriorly; antennomeres I–III distinctly elongate, III longer than II and IV, IV–X successively stouter, moniliform, XI longer than X, narrowed apically.

Pronotum ( Fig. 1B View FIGURE 1 ) about as broad as elytra, transverse, widest at middle, strongly constricted at base; length 0.69– 0.73 mm, width 0.89–0.96 mm; with punctures and setae similar to those on head, with reticulate microsculpture along posterolateral and posterior margins. Mesoscutellum roundly triangular, with reticulate microsculpture.

Elytra ( Fig. 1B View FIGURE 1 ) about as long as wide, length 0.88–0.95 mm, width 0.92–0.93 mm; parallel sided; sparsely punctate, punctures shallower than those of pronotum; basal area lateral to mesoscutellum with reticulate microsculpture.

Legs short; profemora projected ventrally at basal half; protibiae ( Fig. 2A View FIGURE 2 ) curved at basal 1/3, with seven external and two apical spines, with well-developed ctenidium along mesal margin; mesotibia ( Fig. 2B View FIGURE 2 ) with lateral row of macrosetae, with two external and two apical spines; metatibia with apicolateral row of macrosetae, and two apical spines.

Abdomen slightly broadened posteriorly; punctation and setation similar to those on elytra, but sparser and finer, also with dense reticulate microsculpture. Paired tergite IX ( Fig. 2C View FIGURE 2 ) with several exceptionally elongate lateral setae; tergite X ( Fig. 2C View FIGURE 2 ) strongly protruding posterolaterally, with two small denticles at posterior margin. Sternites VII–VIII lacking median impression; sternite VIII ( Fig. 2D View FIGURE 2 ) subtriangular, each side with few long setae and two small, acute denticles; sternite IX ( Fig. 2E View FIGURE 2 ) elongate, weakly sclerotized, apex rounded and finely denticulate.

Aedeagus ( Fig. 2F–G View FIGURE 2 ) elongate, length of median lobe 0.56 mm, length including parameres 0.64 mm; median lobe weakly sclerotized at basal half, apical half strongly curved dorsally and narrowed apically; parameres long, exceeding apex of median lobe, each paramere with two long macrosetae at apex; endophallus with three sclerites, elongate, twisted, and spiral-like.

Female. Unknown.

Distribution. Northern Laos: Houaphanh.

Etymology. We dedicate this new species to Dr. Joerg Dreybrodt ( Germany), a specialist in survey and mapping, and the coordinator of the 2009–2010 survey team.