Aquamegadrili Jamieson, 1988

Aquamegadrili Jamieson, 1988 and Lumbricoidea sensu Jamieson 1971

The Aquamegadrili, named by Jamieson (1988) for aquatic crassiclitellates (see Introduction for geographic distribution), consist of the Sparganophilidae , Biwadrilidae , Almidae (mostly warm tropics, including Criodrilus , Mediterranean region, etc.) and Lutodrilidae (Southern Neartic). In the likelihood ratio tests the 28S data-set is inconsistent with monophyly of the Aquamegadrili but this is entirely due to the inclusion of Komarekiona as sister-taxon of Sparganophilus . The 28S data are inconsistent with partition of crassiclitellates into Aquamegadrili and Terrimegadrili (see families listed in Introduction); instead the representatives ( Sparganophilus , Criodrilus and Lutodrilus ) of the original aquamegadrile taxa, with Komarekiona , lie within a paraphyletic Terrimegadrili (Figs 2; 3). The Lumbricoidea is incompatible with the 28S data but principally because the glossoscolecid sp. never groups with the remainder in the best trees but groups equivocally with Eudrilus . However the unity of the majority of the remaining Lumbricoidea will necessarily require more sampling.