Euconnus (s. str.) glanduliferus (Nietner), 2025

Euconnus (s. str.) glanduliferus (Nietner) , stat. rev.

( Figs 29 View FIGURES 24–32 , 53–54 View FIGURES 53–58 )

Scydmaenus glanduliferus Nietner, 1856: 551 .

Euconnus (Euconnus) glandulifer (Nietner) View in CoL ; Csiki, 1919: 48 (lapsus calami or unjustified emendation of glanduliferus ).

Type material. Lectotype (here designated): ♂, three labels: “ Ceylon / Nietner..s” (white, printed and handwritten, text in black frame], QR code label with collection number 8186, and newly added “ EUCONNUS / (s. str.) / glanduliferus ( Nietner, 1856) / LECTOTYPUS / P. Jałoszyński, 2025 ” ( MIZ). Paralectotype: ♀, with white printed copy of locality label and identification labels with “PARALECTOTYPE” ( MIZ) .

Revised diagnosis. Vertex weakly bulging posterodorsad and with dense bristles; eyes large, in lateral view slightly longer than tempora; antennal clubs tetramerous and longer than remaining flagellomeres combined; pronotum widest at base, and strongly narrowing anterad, with one lateral pair of antebasal pits connected by shallow transverse groove, which mesally reduces in depth and is obliterated at middle; aedeagus in ventral view broadest near middle, drop-shaped, with subtrapezoidal apical region, endophallus symmetrical and composed of complex set of sclerotized elements, parameres nearly reaching apex of median lobe.

Redescription. Body of male ( Fig. 29 View FIGURES 24–32 ) stout, strongly convex, moderately dark brown; setae distinctly lighter than cuticle; BL 1.35 mm.

Head broadest across eyes, HL 0.25 mm, HW 0.28 mm; frontal and anterior vertexal regions confluent, weakly convex, vertexal region weakly bulging posterodorsad; supraantennal tubercles barely marked; eyes large, in lateral view slightly longer than tempora. Punctures on frons and vertex fine and inconspicuous; setae sparse, short and weakly suberect; tempora and vertex with dense and long bristles directed posteriorly. Antennae slender, with sharply delimited tetramerous clubs, AnL 0.55 mm; antennomere 1–2 each weakly elongate, 3–5 each transverse, 6 about as long as broad, 7 transverse, 8–10 each strongly transverse, 11 indistinctly broader than 10, slightly shorter than 9–10 combined, about 1.2 times as long as broad.

Pronotum subtrapezoidal, broadest at base; PL 0.35 mm, PW 0.38 mm; anterior margin weakly arcuate, anterior corners poorly marked, obtuse-angled and blunt; lateral margins only slightly rounded; posterior corners nearly right-angled and blunt; posterior margin shallowly bisinuate; base with two small lateral pits and with shallow transverse groove laterally connecting with pits but mesally becoming shallower and at middle obliterated. Punctures on pronotal disc similar to those on frons and vertex, inconspicuous; setae moderately long, moderately dense and suberect, lateral margins with bristles (but largely broken off in the lectotype).

Elytra together oval, broadest between middle and anterior third; EL 0.75 mm, EW 0.65 mm, EI 1.15; humeral calli distinctly elevated, basal impressions shallow but distinct; elytral apices separately rounded. Punctures and setae similar to those on median region of pronotal disc.

Legs moderately long, slender, unmodified.

Aedeagus ( Figs 53–54 View FIGURES 53–58 ) stout; AeL 0.28 mm; median lobe in ventral view approximately drop-shaped, broadest near middle, narrowing distally and with subtrapezoidal apical region strongly bent dorsad, endophallus symmetrical and composed of complex set of sclerotized elements situated in subapical region, parameres slender, in ventral view slightly recurved, nearly reaching apex of median lobe, each with 3–4 apical and subapical setae.

Female. Externally indistinguishable from male. BL 1.35 mm; HL 0.23 mm, HW 0.28 mm, AnL 0.53 mm; PL 0.38 mm, PW 0.40 mm; EL 0.75 mm, EW 0.63 mm, EI 1.20.

Distribution. Sri Lanka, near Colombo.

Remarks. Three specimens were found in a tray with the label “ glandulifer ”, two conspecific beetles, a male and a female, with tetramerous antennal clubs originally mounted on one card, and a male with trimerous clubs identified here as Euconnus graminicola (Nietner) , mounted separately ( Fig. 27 View FIGURES 24–32 ). Nietner (1856) stated that he collected them “in the sweeping net on the lawns of my garden about sunset.” External features of the two specimens agree with the original description of S. glanduliferus . The male with the tetramerous club is here designated as the lectotype of Scydmaenus glanduliferus Nietner. Franz (1982) does not mention this name (neither E. glandulifer , which was used for this species in Csiki (1919)).

Euconnus glanduliferus belongs to the E. claviger group, which comprises species with subconical pronotum, sharply delimited tetramerous antennal clubs longer than the remaining flagellomeres combined and composed of loosely assembled antennomeres, of which 8–10 are usually transverse, and a drop-shaped aedeagus with strongly developed, and usually complex endophallic sclerites (similar to those in Euconnus claviger ( Müller & Kunze, 1822) ; see Jałoszyński (2015, 2021c)). Species in this group (especially tropical and subtropical ones) are difficult to identify by external characters and primary diagnostic features are those associated with the male genitalia. They all have been included in the subgenus Napochus Thomson, 1859 , later merged with Euconnus s. str. ( Jałoszyński 2021c). Only two species were placed in Napochus in the Sri Lankan monograph by Franz, and their aedeagi were illustrated in dorsal or ventral views, allowing for comparisons. None of them seems to be similar to that of E. glanduliferus .