Leprosoma tuberculatum JAKOVLEV, 1874

Leprosoma tuberculatum JAKOVLEV, 1874

Leprosoma inaequale Horváth 1911 , nov. syn.

( Figs 3, 11-13, 17, 18, 23)

Type material examined ( ZISP). Holotype of L. tuberculatum : ♂, Russia, Astrakhan Prov., “ Bogdo [ Hill ]”, “k. V. Jakovleva ”, “ Leprosoma tuberculatum Jak. V. Jakowlew det.”, “Holotypus ♂ Leprosoma tuberculatum Jak. ” . Lectotype of L. turþestanicum: ♂, Uzbekistan, Karaul-Bazar, 13.IV.1927 (Zolotov) ; paralectotypes: station Ziaddin, 29.V.1927 (Zolotov), 3♀♀ .

Additional material examined ( ZISP). RUSSIA, Astrakhan Prov.: Seroglazinka , 1♂ ; Volgogard Prov.: Elton lake, 5.VI.1961 (Emeljanov & Kerzhner), 1♂, 2♀♀ ; Orenburg Prov.: Elshanka, 24.VII.1933 (Zimin), 2♂ ; ARMENIA: Erivan, 3, 8.VII.1935, 26, 30.VIII.1936 (Richter & Ter-Minasian), 2♂♂, 3♀♀ , 31.X., 1.XI.1931 (Korinek), 1♂, 1♀ , 18.V.1938 (No. 201-938) (Ter-Minasian & Richter), 1♀ ; near Erivan (Stekolnikov), 1♂ ; Dzhervezh, 29.IX.1946 (Richter), 1♂ ; Sovetashen [= Nubarashen], 27.V.1938, 1♀ ; AZERBAIJAN: Nakhichevan, 15.VI.1933 (Znoiko), 1♂ ; Disar near Ordubad, 20.VII.1933 (Znoiko), 1♀ , 2, 5.V.1934 (Ter-Minasian), 3♀ ; Elisavetpol [= Gyandzha ], 24.I.1905, 6.VIII.1906 (Jakovlev coll.), 3♂♂, 4♀♀ ; IRAN: S slopes of Elburz, Tadzhrish, 25-30. VI.1928 (Siyazov), 1♂, 1♀ ; Charbag, 4.V.1914 (Kiritshenko), 2♂♂, 1♀ ; Shakhrud [= Emamshahr ], 23, 26, 27, 29, 31.V, 1-3, 5-7, 11.VI.1914 (Kiritshenko), 12♂♂, 13♀♀ ; Shakhkukh [ Shachkuh superior ], 15, 16, 18-22, 25, 26, 29.VI, 1-5.VII.1914 (Kiritshenko), 22♂♂, 33♀♀ ; Shachkuh-poion, 3-5.VII.1914 (Kiritshenko), 6♂♂, 5♀♀ ; Kazvin, 16.III.1935 (Jenjouriste), 1♀ ; KAZAKHSTAN, West Kazakhstan Prov.: Dzhanybek, 26.VI.1961 (Kerzhner), 1♂, 1♀ ; Kalmykovo, 17.VI.1909 (Uvarov), 1♂, 1♀ ; Atyrau Prov.: Inder lake, 23, 24.VI. 1951 (Steinberg), 1♂ ; UZBEKISTAN: Shirabad, 27-28.V., 2, 4.VII.1912 (Kiritshenko), 12♂♂, 4♀♀ ; Tutak, 31.IX.1911 (Kiritshenko), 1♀ ; Kyzyl-Kum, Ayakguzhumdy , 29.IV.1965 (Emeljanov), 2♂ ; Kyzyl-Kum, 20 km N Ayakuzhumdy , 17.VI.1966 (Kerzhner), 1♂ ; Kyzyl-Kum, 70 km S Tamdy-Bulak , 1.V.1965 (Kerzhner), 1♂, 1 .

Diagnosis. Body moderately large. Humeri broadly or rectangular rounded, long or shortened, directed laterad. Tubercles in anterior portion of pronotum small. Intermediate transverse carina short, slightly curved caudad. Posterior transverse carina strongly sinuous. Tubercles on posterior angles of abdominal segments large. Laterotergites of connexivum each with distinct transverse elevation. Pygophore ( Figs 17, 18) slightly wider than long. Iits ventro-apical margin with shallow, wide, rounded-rectangular notch ( Figs 11, 13) and almost straight, slightly concave lateral parts. Ventral infolding of rim of pygophore very long, concave, perpendicular to ventral wall of pygophore. Parandria with convex, elevated, strongly sclerotized outer margin and straight or concave, membranous inner margin. Outer margin of paramere hypophysis convex just before apex ( Fig. 12). Female genitalia. Caudal margin of first gonocoxae convex near lateral angles and slightly concave near medial angles. Medial part of spermathecal duct extending beyond anterior margin of abdominal sternum V.

Measurements (mm) and ratios. Total length 5.30–6.55, width of pronotum 3.35–4.25, width of abdomen 3.60–4.20, ratio head length/head width 0.98–1.02, ratio scutellum width/scutellum length 0.50–0.55. Length of antennal segments I–V: 12–13: 18–21: 11–13: 11–14: 23–28.

Distribution. Recorded from South of Russian European territory, Asian part of Turkey, Iran, Afghanistan, Armenia, Azerbaijan (new record), Kazakhstan, Uzbekistan, Turkmenistan, Tajikistan, Kyrgyzstan, northwestern China, but some of these records (definitely those from Turkmenistan and Tajikistan) belong to L. tenuimarginatum . Specimens with long humeri are found in Astrakhan and Orenburg Provinces an in west Kazakhstan; specimens with shortened humeri distributed in the rest part of its natural habitat ( Fig. 31).

Biology. The host plants are Brassicaceae .

L. inaequale Horváth (1911) was described from Armenia (Araxis valley). According to the original description, this species is characterised by the following main characters: abdomen wider than pronotum; lateral margins of pronotum distinctly sinuate in anterior part; humeri broadly rounded, slightly protruding forward; base and sides of scutellum and connexivum blackish; total length 6 mm; width of pronotum 3.75 mm; width of abdomen 4 mm. A large series of specimens of L. inaequale from Armenia and Azerbaijan in the collection of ZISP shows a gradual transition between the characters of L. inaequale and L. tuberculatum .The most important character, ratio of width of abdomen/width of pronotum, is presented in Fig. 5. On the basis of these data, L. inaequale is synonymised with L. tuberculatum .

L. turþestanicum Zolotov 1928 was synonymised with L. tuberculatum by Kerzhner (1964). This variety is characterised by the short humeri and short tubercles on posterior angles of abdominal segments and resembles specimens previously placed in L. inaequale .