Copa kabana, Raven, 2015

Copa kabana View in CoL sp. nov.

( Figs 3b View FIGURE 3 , 35 View FIGURE 35 , 36a–e View FIGURE 36 , 135 View FIGURE 135 . Map 17)

Material. Queensland: holotype ♂, Mapleton Falls NP, 26°38'S, 152°51'E, rainforest, flight intercept trap, 30 Nov 1991 – 8 Jan 1992, D.J. Cook, QM S31538 View Materials GoogleMaps . Paratypes: Queensland: 2♂, Boombana NP ( BCC1 View Materials ) , 27°24'S, 152°47'E, rainforest, 7 Nov 2003, QMP, QM S54576 View Materials , QM S65233 View Materials GoogleMaps ; 1♂, Brooyar Fire Tower, via Glastonbury , 26°10'S, 152°28'E, rainforest, pitfall, 23 Aug 1975 – 29 Feb 1976, G. & S. Monteith, QM S29400 View Materials GoogleMaps ; 1♀, Cainbable Ra, Lamington NP, 28°14'S, 153°08'E, rainforest, pitfall, 31 Jan–11 Apr 1976, G. & S. Monteith, QM S29514 View Materials GoogleMaps ; 1♀, Casey Ck, via Imbil , 26°28'S 152°41'E, rainforest, pitfall, 31 Dec 1974 – 27 Mar 1975, G. & S. Monteith, QM S39293 View Materials GoogleMaps ; 1♀, Cooyar, Maidenwell Rd , 26°56'S, 151°49'E, rainforest, pitfall, 25 Jan–1 May 1976, G. & S. Monteith, QM S29406 View Materials GoogleMaps ; 1♀, Cunninghams Gap , 28°03'S, 152°24'E, rainforest, intercept flight trap, 1 Dec 1991 – 6 Jan 1992, D.J. Cook, QM S39295 View Materials GoogleMaps ; 1♀, Curtis Farm, Canungra , 28°04'S, 153°07'E, rainforest, pitfall, 27 Nov 1976 – 23 Jan 1977, G. & S. Monteith, QM S29401 View Materials GoogleMaps ; 1♀, Dwyer Creek, Imbil , 26°28'S, 152°38'E, hoop pine scrub, flight intercept trap, 24 Sep 2001 – 15 Jan 2002, G. Monteith, QM S60632 View Materials GoogleMaps ; 1♂ 1♀, Kroombit Tops (site 5), 24°25'S, 151°03'E, rainforest, pitfall, 10–18 Dec 1983, G. Monteith, V GoogleMaps . Davies, J. Gallon, G. Thompson, QM S29476 View Materials ; 1♀, Kroombit Tops, Beauty Spot 98, 24°22'S, 151°01'E, rainforest, 9–19 Dec 1983, G. Monteith, G. Thompson, QM S29494 View Materials GoogleMaps ; 1♀, Kroombit Tops, Beauty Spot 98 (site 2), 24°25'S, 151°03'E, rainforest, pitfall, 10–18 Dec 1983, G. Monteith, V GoogleMaps . Davies, J. Gallon, G. Thompson, QM S29496 View Materials ; 1♀, Lamington NP, Lukes Bluff Rd , 28°19'S, 153°05'E, rainforest, pitfall, 12–27 Dec 1981, G. Monteith, R GoogleMaps . Raven, D. Yeates, QM S29588 View Materials ; 1♀, Lamington NP, O'Reillys, Wishing Tree Circuit , 28°15'S, 153°09'E, rainforest, pitfall, 9 Feb 1982 – 19 Mar 1982, J. Gallon, D. Yeates, G. Thompson, QM S29487 View Materials GoogleMaps ; 1♂, Little Yabba Ck , 26°37'S, 152°41'E, rainforest, intercept flight trap, 30 Nov 1991 – 7 Jan 1992, D.J. Cook, QM S25199 View Materials GoogleMaps . Mapleton Falls NP, 26°38'S, 152°51'E, rainforest, intercept flight trap, D.J. Cook: 1♂, 8 Jan–3 Mar 1992, QM S52242 GoogleMaps ; allotype ♀, 30 Nov 1991 – 8 Jan 1992, QM S31539 View Materials ; 1♂ 1♀, same data, QM S31540 View Materials . 1♂ 1♀, Mary Cairncross Pk , 26°46'S, 152°52'E, rainforest, flight intercept trap, 29 Nov 1991 – 8 Jan 1992, D.J. Cook, QM S43405 View Materials GoogleMaps ; 1♂, Mooloolah R, (Glenview) , 26°45'S, 153°00'E, rainforest, pitfall, 27 Sep 1977 – 2 Jan 1978, G. & S. Monteith, QM S29405 View Materials GoogleMaps ; 1♀, Mt Archer, Kilcoy , 26°57'S 152°34'E, rainforest, 29 Apr 1985, J. Gallon, QM S29385 View Materials GoogleMaps . Mt Glorious , 27°20'S, 152°46'E, rainforest, malaise trap, A. Hiller: 1♂, 10 Aug–13 Nov 1983, QM S29500 View Materials GoogleMaps ; 1♂, Sep 1978, QM S39291 GoogleMaps ; 1♂, Mt Glorious , 27°20'S, 152°46'E, malaise trap, Jun–Jul 1977, QM S29386 View Materials GoogleMaps ; ♀, 28 Feb 1990, K. Hiller, QM S15655 View Materials ; 1♂ 1♀, Mt Goonaneman , 25°26'S, 152°08'E, pitfall, 3–7 Nov 1980, R GoogleMaps . Raven , V . Davies , QM S29383 View Materials ; 1♂, Mt Mee , 27°03'S, 152°41'E, rainforest, intercept flight trap, 29 Nov 1991 – 8 Jan 1992, D.J. Cook, QM S31638 View Materials GoogleMaps ; 1♂ 1♀, Mt Tamborine , 27°58'S, 153°12'E, rainforest, pitfall, Dec 1978 – Jan 1979, J. Grimshaw, QM S29384 View Materials GoogleMaps ; 1♂ 1♀, Nichols Scrub , 28°11'S, 153°26'E, rainforest, intercept flight trap, 1 Jan 1992 – 4 Mar 1992, D.J. Cook, QM S25072 View Materials GoogleMaps ; 1♀, Nichols Scrub , 28°11'S, 153°26'E, rainforest, intercept flight trap, 9 Dec 1991 – 1 Jan 1992, D.J. Cook, QM S25069 View Materials GoogleMaps . Ravensbourne NP, 27°22'S, 152°11'E, rainforest, pitfall: 1♂, 7 Jan–1 Mar 1992, D.J. Cook, QM S31639 View Materials GoogleMaps ; 1♀, 1 Dec 1991 – 7 Jan 1992, QM S25063 View Materials ; 1♀, intercept flight trap, 7 Jan–1 Mar 1992, QM S25170 View Materials . 1♂, Saddletree Ck, Bunya Mtns , 26°51'S, 151°38'E, rainforest, pitfall, 1 Dec 1991 – 7 Jan 1992, D.J. Cook, QM S50418 View Materials GoogleMaps ; 1♀, Sunday Ck , 26°40'S, 152°34'E, rainforest, pitfall, 18 Dec 1996 – 20 Jan 1997, G. Monteith, QM S39292 View Materials GoogleMaps ; 1♀, The Head, via Killarney , 28°23'S, 152°19'E, rainforest, pitfall, 17 Nov–27 Dec 1974, G. & S. Monteith, QM S39294 View Materials GoogleMaps ; 1♀, Toowoomba, Boyce Reserve ( GMP60 ) , 27°34'S, 151°57'E, pitfall, 1976–1977, G. & S. Monteith, QM S29583 View Materials GoogleMaps ; 1♀, Tungi Ck, via Jimna , 26°40'S, 152°28'E, rainforest, intercept flight trap, 18 Dec 1996 – 20 Jan 1997, G. Monteith, QM S39296 View Materials GoogleMaps ; 1♂, Tungi Ck, via Jimna , 26°40'S, 152°28'E, rainforest, pitfall, 18 Dec 1996 – 20 Jan 1997, G. Monteith, QM S39290 View Materials GoogleMaps ; 1♀, Upper Tallebudgera Valley , 28°15'S, 153°16'E, rainforest, pitfall, 8 Jan–17 Mar 1985, G. Monteith, G. Thompson, D. Cook, QM S29478 View Materials GoogleMaps ; 1♀, Enterprise Mine, Mallee #2, 27°35'S, 153°26'E, mallee, 7 Jan 2002, QMP, QM S56290 View Materials GoogleMaps ; 1♂ 1♀, Little Yabba Ck , 26°37'S, 152°41'E, rainforest, intercept flight trap, 7 Jan–2 Mar 1992, D.J. Cook, QM S43400 View Materials GoogleMaps ; 1♀, Yarraman, Cooyar Ra , 26°52'S, 151°54'E, rainforest, pitfall, 1 Dec 1991 – 7 Jan 1992, D.J. Cook, QM S43402 View Materials GoogleMaps . New South Wales. 1♂, Bruxner Park , 30°14'S, 153°06'E, rainforest, 12 Nov 1980, R GoogleMaps . J. Raven , QM S29511 View Materials ; 1♀, Enterprise Mine, Mallee #2, 27°35'S, 153°26'E, mallee, 7 Jan 2002, QMP, QM S56290 View Materials GoogleMaps ; 1♂ 1♀, Little Yabba Ck , 26°37'S, 152°41'E, rainforest, intercept flight trap, 7 Jan–2 Mar 1992, D.J. Cook, QM S43400 View Materials GoogleMaps ; 1♀, Yarraman, Cooyar Ra , 26°52'S, 151°54'E, rainforest, pitfall, 1 Dec 1991 – 7 Jan 1992, D.J. Cook, QM S43402 View Materials GoogleMaps .

Diagnosis. Males differ from those of other Australian Castianeirinae in the absence of a paracymbial spine and cylindrical male palpal tibia, and females differ from those of Ozcopa , which they closely resemble, in the much smaller copulatory fossae. Males of Copa kabana sp. nov. differ from those species in the sinuous, not spiralled embolus ( Fig. 36b View FIGURE 36 ) and females in the shorter copulatory ducts ( Fig. 36e View FIGURE 36 ).

Etymology. The species epithet is a phonetic version of the famous beach in Brasil, Copacabana.

Description holotype male QM S31538 View Materials

Carapace 3.52 long, 2.64 wide. Abdomen 2.96 long, 1.92 wide. Total length 6.5.

Colour: carapace orange brown with dark brown scallops along edges and band of brown laterally and on clypeus (darkest in posterior third); legs orange brown, femora darker with brown areas distally on femora I–III, tibiae III and IV, and in basal half of metatarsi IV; a bdomen brown dorsally with central pallid region, narrowing posteriorly, ventrally dark brown with narrow pallid flanks. Sternum, maxillae and coxae light orange brown. Carapace: light cover of pallid and black feathery hairs with few long bristles; margins with evenly sparse horizontally directed short black setae; fovea straight, closed. Lateral profile of carapace ( Fig. 3b View FIGURE 3 ): high but more or less flat or slightly arched for its length; marginally highest point behind fovea at shallowly sloping posterior declivity; eyes set behind clypeus edge. Retrolateral face of large chelicerae smooth. Eyes: AME largest; with pile of white hairs lateral of group. Chelicerae: vertical and deep (front to back); fang shield small rounded. Fang slightly diagonal, groove long, almost vertical; two teeth high on retromargin and one large and one small basally on promargin plus very small tooth distally. Retrobasal ridge low, distinct, setose. Maxillae: serrula long; inner margin excavate, sloping, outer convex; anterior edge truncate. Labium ca. as long as wide. Sternum widely cordate, intercoxal projections on I/II, II/III; separate pre-coxal sclerites at II–IV; uniformly and lightly setose. Legs: trochanters clearly notched, ca. twice as wide as deep; light to very light cover of dark feathery hairs on femora to metatarsi; erect scopula hairs sparse on metatarsi and tarsi I and II. Spines: (erect) I: fe pv1p2d3r4; pa 0; ti v2.2.2; me v2.2. II: fe pv1p2d3r3; pa 0; ti v2.2.2; me v2.2. III: fe p3d3r3; pa 0; ti p2d1r2v2.2.2; me p3r3v2.2.2. IV: fe p3d3r2; pa 0; p2r2v2.2.2; me p3r3v2.2.2. Palp: fe p1d2; rest long bristles. Claws: with ca. 5 teeth with tufts similar on I and IV. Abdomen: covered with feathery hairs. Scutes: dorsal small, oval, lightly sclerotised; genital scute without anterior collar or thickened edge; tracheal spiracle overhung by small triangular scute with posterior nipple. Pedicel a small winged triangle from below, not fused to sternum. Spinnerets: PLS about 2/3 size of ALS and clearly bigger than PMS. Palp: tibia cylindrical, save for shallow ventral depression for length; cymbium long, without paracymbial spine, symmetrical groove to tip, without apical spine or scopula; large trianguloid scooped plate is part of basal haematodocha; bulb simple, narrow pyriform with slight enlargement at embolus base, but then tapering gradually in long fine gently curved hook; sperm duct large, overlaps itself basally and mid-distally, then undulates into embolus; prolateral cymbium basally with depression locking with opposed tibial process.

Allotype female QM S31539 View Materials . Like ♂ except as follows:

Carapace 3.44 long, 2.48 wide. Abdomen 4.24 long, 2.80 wide. Total length 7.7.

Chelicerae with one large and one small basal tooth on promargin, two large on retromargin; fang shield distinct, squared with many long thick curved bristles; retrobasally with band of short thorn-like setae. Spines: on tibiae I and II and metatarsi, ventral spines long strong and overlapping (5–10%); other leg spines also long, strong. I: fe pv1p2d3r1; ti v2.2.2; me v2.2. II: fe pv1p2d3r2; ti v2.2.2; me v2.2. III: fe p3d3r3; pa 1 apical bristle; ti p2d1r2v.2.2.0; me p3r3v2.2.3. IV: fe p2d3r2; pa 1 apical bristle; ti p2d1r2v1.2.2; me p3r3v2.2.3(1). Palp: fe p1d2; pa p1d2; ti p2 basal d1; ta p2r1, all basal, v3 apically. Legs: feathery hairs on femora–metatarsi; no scopula. Claw tufts dense, similar on I and IV. 5–7 teeth on paired claws. Palp: slender curved claw with ca. 3 small teeth; tarsus converges gently to tip without dense hairs of pre-apical widening. Abdomen: with small square scute on anterior face. Spinnerets: PMS large with three enlarged cylindrical gland spigots apically; PLS slightly smaller than ALS with 2 large cylindrical gland spigots, ALS with ental subfield of small pyriform gland spigots. Tracheal scute like ♂. No dorsal or ventral scute, weakly sclerotised genital scute. Epigyne: broad with two very small lateral fossae directed medially, internally with copulatory ducts and ridges directed medially, large anterior lobe and beanshaped smaller posterior lobe.

Distribution and Habitat. Widespread in rainforests of south-eastern Queensland to northern New South Wales, from Kroombit Tops , through the Conondale and D'Aguilar Ranges to the Border Ranges.

MAPS 13–18. Records of Australian species of Battalus Karsch, 1878 , Copa Simon, 1985 , and Disnyssus gen. nov.